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nipplebract arrowhead

arrowhead family, water-plantain family

Habit Herbs, perennial, to 112 cm; rhizomes not coarse; stolons absent; corms absent. Herbs, annual or perennial, rhizomatous, stoloniferous, or cormose, caulescent, glabrous to stellate-pubescent; sap milky.
Roots

septate or not septate.

Leaves

submersed and emergent;

petiole triangular, 11.5–35 cm;

blade elliptic to ovate, 25–56 cm.

basal, submersed, floating, or emersed, sessile or petiolate, sheathing proximally;

blade with translucent markings of dots or lines present or absent, basal lobes present or absent;

venation reticulate, primary veins parallel from base of blade to apex, secondary veins reticulate.

Inflorescences

racemes or panicles, of 4–10 whorls, emersed, 8–25 × 5–36 cm;

peduncles to 76 cm;

bracts connate ¼ total length, lanceolate, 4–8 mm, papillose; fruiting pedicels ascending to spreading, cylindric, 1–4.5 cm.

scapose racemes or panicles, rarely umbels, erect, rarely floating or decumbent, whorled (forming racemes) or whorls branching (forming panicles), bracteolate.

Flowers

to 3 cm diam.;

sepals appressed to spreading;

filaments slightly dilated, shorter than anthers, glabrous;

pistillate pedicellate, without ring of sterile stamens.

bisexual or unisexual, if unisexual, staminate and pistillate on same or different plants, hypogynous, subsessile to long-pedicellate;

sepals persistent, 3;

petals deciduous, 3, delicate;

stamens 0, 6, 9, or to 30, distinct;

anthers 2-loculed, dehiscing longitudinally;

pistils 0 or 6–1500 or more, distinct or coherent proximally, 1-loculed;

placentation basal;

ovules1–2.

Fruits

achenes or follicles.

Seeds

embryo U-shaped;

endosperm absent in mature seed.

Fruiting

heads 0.7–1 cm diam.;

achenes obovoid, abaxially keeled, 1.2–1.5 × 0.7–1.1 mm, beaked;

faces not tuberculate, wings 1, ± entire, glands 0–1;

beak lateral, erect, 0.1–0.3 mm.

Sagittaria papillosa

Alismataceae

Phenology Flowering spring–fall.
Habitat Swamps, marshes, bogs, ditches, borders of lakes
Elevation 0–300 m (0–1000 ft)
Distribution
from FNA
AR; LA; OK; TX
[BONAP county map]
Nearly worldwide; primarily tropical and subtropical regions
[BONAP county map]
Discussion

Genera 12, species ca. 80 (4 genera, 34 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistils weakly coherent proximally into starlike aggregation; petals erose
Damasonium
1. Pistils distinct, forming heads or rings; petals entire.
→ 2
2. Pistils arranged in ring around margin of flattened receptacle.
Alisma
2. Pistils spirally arranged on convex receptacle.
→ 3
3. Flowers all bisexual; fruits mostly plump, longitudinally ribbed, lateral wings absent.
Echinodorus
3. Flowers unisexual (at least the proximal); fruits compressed, lateral wing often present, 1, curved.
Sagittaria
Source FNA vol. 22. FNA vol. 22, p. 7. Authors: Robert R. Haynes, C. Barre Hellquist.
Parent taxa Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
Alisma, Damasonium, Echinodorus, Sagittaria
Name authority Buchenau: Index Crit. Butom. Alism. Juncagin. 27, 44. (1868) Ventenat
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