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annual sagittaria, California arrowhead, California lophotocarpus, giant arrowhead, hooded arrowhead, sagittaire spongieuse, spongy-leaf arrowhead

arrowhead family, water-plantain family

Habit Herbs, annual or perennial, to 100 cm; rhizomes present; stolons present; corms present. Herbs, annual or perennial, rhizomatous, stoloniferous, or cormose, caulescent, glabrous to stellate-pubescent; sap milky.
Roots

septate or not septate.

Leaves

submersed and emersed; submersed leaves absent or phyllodial, flattened, to 17 × 2 cm; emersed leaves with petiole triangular, 21–55 cm, blade hastate to sagittate, 2.5–17.5 × 0.6–22 cm, lobes longer than or equal to remainder of blade.

basal, submersed, floating, or emersed, sessile or petiolate, sheathing proximally;

blade with translucent markings of dots or lines present or absent, basal lobes present or absent;

venation reticulate, primary veins parallel from base of blade to apex, secondary veins reticulate.

Inflorescences

racemes or panicles, of 1–15 whorls, floating or emersed, 1.5–28 × 1.5–15 cm;

peduncles 15–47 cm;

bracts distinct or connate less than ¼ total length, lanceolate to elliptic, 4–34 mm, delicate, not papillose; fruiting pedicels recurved, club-shaped, 0.5–4.2 cm.

scapose racemes or panicles, rarely umbels, erect, rarely floating or decumbent, whorled (forming racemes) or whorls branching (forming panicles), bracteolate.

Flowers

2–5 cm diam.;

sepals spreading in staminate, erect in pistillate, enclosing flower or fruiting head;

filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, with or without ring of sterile stamens.

bisexual or unisexual, if unisexual, staminate and pistillate on same or different plants, hypogynous, subsessile to long-pedicellate;

sepals persistent, 3;

petals deciduous, 3, delicate;

stamens 0, 6, 9, or to 30, distinct;

anthers 2-loculed, dehiscing longitudinally;

pistils 0 or 6–1500 or more, distinct or coherent proximally, 1-loculed;

placentation basal;

ovules1–2.

Fruits

achenes or follicles.

Seeds

embryo U-shaped;

endosperm absent in mature seed.

Fruiting

heads 1.2–2.1 cm diam.;

achenes oblanceoloid, not abaxially keeled, 2–4.3 × 0.7–1.5 mm, beaked;

faces not tuberculate, wings absent, glands 1;

beak lateral, horizontal, 0.4–0.8 mm.

Sagittaria montevidensis

Alismataceae

Distribution
from FNA
AL; AR; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; SC; SD; TN; TX; VA; WI; WV; NB; Mexico; South America
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Nearly worldwide; primarily tropical and subtropical regions
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Discussion

Subspecies 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 12, species ca. 80 (4 genera, 34 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants of tidal mud flats, often stranded by low tide, emersed blades spatulate, rarely sagittate.
subsp. spongiosa
1. Plants of fresh waters, not stranded by low tide; emersed blades hastate to sagittate.
→ 2
2. Pistillate flowers without ring of sterile stamens; petals with purple spot at base
subsp. montevidensis
2. Pistillate flowers with ring of sterile stamens; petals without purple spot at base.
subsp. calycina
1. Pistils weakly coherent proximally into starlike aggregation; petals erose
Damasonium
1. Pistils distinct, forming heads or rings; petals entire.
→ 2
2. Pistils arranged in ring around margin of flattened receptacle.
Alisma
2. Pistils spirally arranged on convex receptacle.
→ 3
3. Flowers all bisexual; fruits mostly plump, longitudinally ribbed, lateral wings absent.
Echinodorus
3. Flowers unisexual (at least the proximal); fruits compressed, lateral wing often present, 1, curved.
Sagittaria
Source FNA vol. 22. FNA vol. 22, p. 7. Authors: Robert R. Haynes, C. Barre Hellquist.
Parent taxa Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
S. montevidensis subsp. calycina, S. montevidensis subsp. montevidensis, S. montevidensis subsp. spongiosa
Alisma, Damasonium, Echinodorus, Sagittaria
Name authority Chamisso & Schlechtendal: Linnaea 2: 156. (1827) Ventenat
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