Sagittaria isoetiformis |
Sagittaria montevidensis |
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quillwort arrowhead |
annual sagittaria, California arrowhead, California lophotocarpus, giant arrowhead, hooded arrowhead, sagittaire spongieuse, spongy-leaf arrowhead |
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Habit | Herbs, perennial, to 65 cm; rhizomes absent; stolons present; corms present. | Herbs, annual or perennial, to 100 cm; rhizomes present; stolons present; corms present. | ||||||||
Leaves | emersed or submersed, phyllodial or rarely dilated apically, flattened, 4–40 cm × 0.05–0.4(–0.5) cm. |
submersed and emersed; submersed leaves absent or phyllodial, flattened, to 17 × 2 cm; emersed leaves with petiole triangular, 21–55 cm, blade hastate to sagittate, 2.5–17.5 × 0.6–22 cm, lobes longer than or equal to remainder of blade. |
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Inflorescences | racemes, of 1–5 whorls, emersed, 2–20 × 1.5–12 cm; peduncles 7–59 cm; bracts connate more than or equal to ¼ total length, oblanceolate, 0.2–0.3 mm, delicate, not papillose; fruiting pedicels spreading, cylindric, 0.5–6 cm. |
racemes or panicles, of 1–15 whorls, floating or emersed, 1.5–28 × 1.5–15 cm; peduncles 15–47 cm; bracts distinct or connate less than ¼ total length, lanceolate to elliptic, 4–34 mm, delicate, not papillose; fruiting pedicels recurved, club-shaped, 0.5–4.2 cm. |
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Flowers | to 1.3 cm diam.; sepals recurved to spreading, not enclosing flower; filaments dilated, shorter than anthers, minutely tomentose; pistillate pedicellate, without ring of sterile stamens. |
2–5 cm diam.; sepals spreading in staminate, erect in pistillate, enclosing flower or fruiting head; filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, with or without ring of sterile stamens. |
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Fruiting | heads 0.5–1 cm diam.; achenes obovoid, abaxially keeled, 2.2–2.8 × 1.5–2 mm, beaked; faces not tuberculate, wings 1–2, ± entire, glands 3–5; beak lateral, incurved-erect, 0.2 mm. |
heads 1.2–2.1 cm diam.; achenes oblanceoloid, not abaxially keeled, 2–4.3 × 0.7–1.5 mm, beaked; faces not tuberculate, wings absent, glands 1; beak lateral, horizontal, 0.4–0.8 mm. |
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Sagittaria isoetiformis |
Sagittaria montevidensis |
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Phenology | Flowering summer–fall. | |||||||||
Habitat | Shores of sandy-bottomed lakes, se coastal plain | |||||||||
Elevation | 0–100 m (0–300 ft) | |||||||||
Distribution |
AL; FL; GA; NC; SC |
AL; AR; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; SC; SD; TN; TX; VA; WI; WV; NB; Mexico; South America
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Discussion | Sagittaria isoetiformis has often been misidentified as Sagittaria teres (E. O. Beal 1960b). The two species can be separated, however: S. teres has nearly terete phyllodia, whereas S. isoetiformis has flattened phyllodia (R. K. Godfrey and P. Adams 1964). A study of the genetics of the two species shows them to be genetically different, and the data indicate the two taxa actually should be considered at the specific level. The two species are capable of CAM photosynthesis, a process very uncommon in the genus and found among none of their supposedly closely related species (A. L. Edwards, pers. comm.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 22. | FNA vol. 22. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | J. G. Smith: Rep. (Annual) Missouri Bot. Gard. 6:115, plate 53. (1895) | Chamisso & Schlechtendal: Linnaea 2: 156. (1827) | ||||||||
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