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guyanese arrowhead

delta arrowhead

Habit Herbs, perennial, to 150 cm; rhizomes absent; stolons present; corms present.
Leaves

submersed and emersed; submersed sessile, phyllodial, flattened, to 26 × 0.5 cm; emersed with petiole ± triangular, 21–70.5 cm, blade linear-ovate to ovate, 4.6–16.4 × 0.7–6.1 cm.

Inflorescences

racemes, of 3–9 whorls, emersed, 2.5–10 × 2–4.5 cm;

peduncles 22–60 cm;

bracts connate more than ¼ total length, lanceolate, 3–5.5 mm, delicate, not papillose; fruiting pedicels spreading to recurved, cylindric, 0.5–3 cm.

Flowers

to 1.8 cm diam.;

sepals spreading to recurved, not enclosing flower or fruiting head;

filaments dilated, longer than anthers, pubescent;

pistillate pedicellate, without ring of sterile stamens.

Fruiting

heads 0.7–1.2 cm diam.;

achenes oblanceoloid, not abaxially keeled, 1.2–2 × 0.8–1.2 mm, beaked;

faces tuberculate, wings absent, glands absent;

beak lateral, horizontal to erect, 0.3–0.6 mm.

2n

= 22.

Sagittaria guayanensis

Sagittaria platyphylla

Phenology Flowering summer–fall.
Habitat Streams and lakes
Elevation 0–900 m (0–3000 ft)
Distribution
from FNA
LA; Mexico; West Indies; tropical regions; Asia; Africa [Introduced, North America]
[BONAP county map]
from FNA
AL; AR; FL; GA; IL; KY; LA; MO; MS; OH; OK; PA; TN; TX; WV; Mexico (Nuevo León Mexico, sw Mexico); Central America (Panama)
[WildflowerSearch map]
[BONAP county map]
Discussion

The name has often been spelled guyanensis (K. Rataj 1972), which is incorrect. Sagittaria guayanensis was the spelling in the protologue. The holotype was collected by Humboldt and Bonpland in Colombia, not Guyana. The type citation is Colombia: Guainia: in wetlands near the sugar mill of Don Felix Farreras and the city of Bolivar. The type is supposedly at Paris, as with all HBK types; however, we (with the help of Alicia Lourteig) have been unable to locate it. MO has a fragment, however. Rataj did designate a neotype: Suriname, Hostman 870 (TCD!). The fragment at MO takes precedence over this neotype.

Sagittaria guayanensis was divided into two subspecies, S. guayanensis subsp. guayanensis from the Neotropics and subsp. lappula (D. Don) Bogin from the Paleotropics. The two subspecies were separated by the shape and size of the fruit in addition to their distribution. Sagittaria guayanensis subsp. lappula has compressed achenes longer than 2.5 mm, whereas subsp. guayanensis has plump achenes shorter than 2.5 mm (C. Bogin 1955).

Subspecies 2 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sagittaria platyphylla has been accepted at the variety level, i.e., Sagittaria graminea var. platyphylla (C. Bogin 1955; J. W. Wooten 1973; E. O. Beal et al. 1982). After studying dozens of populations in the field from much of its range and hundreds of herbarium specimens, we have concluded that this taxon should be recognized at the specific level instead, a conclusion supported by cladistic analyses of morphologic characters (R. M. Kortright 1998)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 22. FNA vol. 22.
Parent taxa Alismataceae > Sagittaria Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
S. guayanensis subsp. guayanensis
Synonyms S. graminea var. platyphylla, S.. mohrii
Name authority Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 1: 250. (1816) (Engelmann) J. G. Smith: N. Amer. Sagittaria. 29. (1894)
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