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grass-leaf arrowhead, grassy arrowhead, sagittaire a feuilles de graminees

awl-leaf arrowhead

Habit Herbs, perennial, to 100 cm; rhizomes coarse; stolons absent; corms absent. Herbs, perennial, to 40 cm, on tidal muds; rhizomes absent; stolons present; corms present.
Leaves

submersed or emersed; submersed leaves phyllodial, angled abaxially, flattened adaxially, 6.4–35 × 0.5–4 cm; emersed with petiole triangular, 6.5–17 cm, blade linear to linear-oblanceolate, 2.5–17.4 × 0.2–4 cm.

submersed, phyllodial, lenticular, 5–40 ´ 0.1–0.4 cm, or rarely floating;

petiole terete, 2.4–4 cm;

blade linear-lanceolate to ovate, 1–2.5 ´ 0.3–1.5 cm.

Inflorescences

racemes or panicles, of 1–12 whorls, emersed, 2.5–21 × 1–8 cm;

peduncles 6.5–29.7 cm;

bracts connate more than to equal to ¼ total length, broadly subulate to lanceolate, 20–50 mm, coarse, not papillose; fruiting pedicels spreading, cylindric, 0.5–5 cm.

racemes, of 2–7 whorls, floating, 2–11 ´ 1.5–4.5 cm;

peduncles 5–40 cm;

bracts connate more than ¼ total length, subulate, 1.5–4.2 mm, delicate, not papillose; fruiting pedicels recurved, cylindric to club-shaped, 0.2–1.1 cm.

Flowers

to 2.3 cm diam.;

sepals recurved to spreading, not enclosing flower;

filaments dilated, shorter than anthers, pubescent;

pistillate flowers pedicellate, without ring of sterile stamens.

0.4–1.2 cm diam.;

sepals spreading to recurved in staminate, erect in pistillate, enclosing flower or fruiting head;

filaments dilated, longer than anther, glabrous;

pistillate pedicellate, occasionally with ring of sterile stamens.

Fruiting

heads 0.6–1.5 cm diam.;

achenes oblanceoloid, without abaxial keel, 1.5–2.8 × 1.1–1.5 mm, beaked;

faces not tuberculate, abaxial wings 0–1, ± entire, glands 1–2;

beak lateral, erect, 0.2 mm.

heads 0.55–0.8 cm diam.;

achenes oblanceoloid, abaxially keeled, 2 ´ 1.5 mm, beaked;

faces not tuberculate, wings 1–2, crenate, glands 0–1;

beak lateral, erect, 0.2–0.4 mm.

2n

= 22.

Sagittaria graminea

Sagittaria subulata

Phenology Flowering summer–fall.
Habitat Streams and brackish bays
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; NB; NF; NS; ON; PE; QC; West Indies (Cuba)
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[BONAP county map]
from FNA
AL; CT; DC; DE; FL; GA; MA; MD; MS; NC; NJ; NY; PA; RI; SC; VA; South America (Colombia)
[WildflowerSearch map]
[BONAP county map]
Discussion

Seven varieties of Sagittaria graminea have been recognized, i.e., var. graminea, var. platyphylla Engelmann, var. teres (S. Watson) Bogin, var. weatherbiana Fernald, var. cristata (Engelmann) Bogin, var. chapmanii J. G. Smith, and var. macrocarpa (J. G. Smith) Bogin (C. Bogin 1955). We accept only one infraspecific rank, i.e., subspecies. Consequently, we have made the appropriate combinations. We accept all of the taxa accepted by Bogin at the varietal level. At specific level we accept Bogin’s var. platyphylla, var. teres, and var. cristata and at subspecific level his var. graminea, var. chapmanii, and var. weatherbiana.

Sagittaria graminea var. macrocarpa actually is synonymous with var. graminea (E. O. Beal 1960b). We therefore are following Beal in recognizing var. macrocarpa sensu Bogin as S. fasciculata. We also accept var. platyphylla, var. teres, and var. cristata at the specific level, leaving only three subspecies. These subspecies can be separated by the branching of the inflorescence and the length of pistillate pedicels.

Subspecies 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sagittaria subulata occurs in shallow brackish waters near the coast. The plants are especially common in areas that are exposed during low tides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences panicles.
subsp. chapmanii
1. Inflorescences racemes.
→ 2
2. Pistillate pedicels 0.5–3 cm; phyllodia less than 1 cm wide
subsp. graminea
2. Pistillate pedicels 2.1–5 cm; phyllodia more than 1 cm wide
subsp. weatherbiana
Source FNA vol. 22. FNA vol. 22.
Parent taxa Alismataceae > Sagittaria Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. teres
Subordinate taxa
S. graminea subsp. chapmanii, S. graminea subsp. graminea, S. graminea subsp. weatherbiana
Synonyms Alisma subulatum, S. lorata, S. subulata var. natans
Name authority Michaux: Flora Boreali-Americana 2: 190. (1803) (Linnaeus) Buchenau: Abhandlungen herausgegeben vom naturwissenschaftlichen Vereine zu Bremen 2: 490. (1871)
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