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grass-leaf arrowhead, grassy arrowhead, sagittaire a feuilles de graminees

Appalachian arrowhead, longbeak arrowhead

Habit Herbs, perennial, to 100 cm; rhizomes coarse; stolons absent; corms absent. Herbs, perennial, to 130 cm; rhizomes absent; stolons present; corms present.
Leaves

submersed or emersed; submersed leaves phyllodial, angled abaxially, flattened adaxially, 6.4–35 × 0.5–4 cm; emersed with petiole triangular, 6.5–17 cm, blade linear to linear-oblanceolate, 2.5–17.4 × 0.2–4 cm.

emersed;

petiole 5-winged, 19–85 cm;

blade sagittate, 3–19 × 2.5–11 cm, basal lobes ± equal to remainder of blade.

Inflorescences

racemes or panicles, of 1–12 whorls, emersed, 2.5–21 × 1–8 cm;

peduncles 6.5–29.7 cm;

bracts connate more than to equal to ¼ total length, broadly subulate to lanceolate, 20–50 mm, coarse, not papillose; fruiting pedicels spreading, cylindric, 0.5–5 cm.

racemes, of 5–12 whorls, emersed, 10–29 × 3–5 cm;

peduncles 25–105 cm;

bracts distinct or if connate, then less than ¼ total length, lanceolate, 7–30 mm, papery, not papillose; fruiting pedicels spreading to ascending, cylindric, 0.3–2.3 cm.

Flowers

to 2.3 cm diam.;

sepals recurved to spreading, not enclosing flower;

filaments dilated, shorter than anthers, pubescent;

pistillate flowers pedicellate, without ring of sterile stamens.

to 3 cm diam.;

sepals recurved to spreading, not enclosing flower or fruiting head;

filaments cylindric, longer than anthers, glabrous;

pistillate pedicellate, without ring of sterile stamens.

Fruiting

heads 0.6–1.5 cm diam.;

achenes oblanceoloid, without abaxial keel, 1.5–2.8 × 1.1–1.5 mm, beaked;

faces not tuberculate, abaxial wings 0–1, ± entire, glands 1–2;

beak lateral, erect, 0.2 mm.

heads 1–2.2 cm diam.;

achenes obovoid, without abaxial keel, 2.1–3.2 × 1.4–2.3 mm, beaked;

faces not tuberculate, wings 0–2, ± entire, glands absent;

beak lateral, strongly recurved, 4–17 mm.

2n

= 22.

Sagittaria graminea

Sagittaria australis

Phenology Flowering summer–early fall (Jul–Oct).
Habitat Slightly basic to slightly acidic ponds, lakes, and swamps
Elevation 1–300 m (0–1000 ft)
Distribution
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; NB; NF; NS; ON; PE; QC; West Indies (Cuba)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; IA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Seven varieties of Sagittaria graminea have been recognized, i.e., var. graminea, var. platyphylla Engelmann, var. teres (S. Watson) Bogin, var. weatherbiana Fernald, var. cristata (Engelmann) Bogin, var. chapmanii J. G. Smith, and var. macrocarpa (J. G. Smith) Bogin (C. Bogin 1955). We accept only one infraspecific rank, i.e., subspecies. Consequently, we have made the appropriate combinations. We accept all of the taxa accepted by Bogin at the varietal level. At specific level we accept Bogin’s var. platyphylla, var. teres, and var. cristata and at subspecific level his var. graminea, var. chapmanii, and var. weatherbiana.

Sagittaria graminea var. macrocarpa actually is synonymous with var. graminea (E. O. Beal 1960b). We therefore are following Beal in recognizing var. macrocarpa sensu Bogin as S. fasciculata. We also accept var. platyphylla, var. teres, and var. cristata at the specific level, leaving only three subspecies. These subspecies can be separated by the branching of the inflorescence and the length of pistillate pedicels.

Subspecies 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Sagittaria longirostra (Micheli) J. G. Smith has been misapplied to S. australis (J. G. Smith) Small (E. O. Beal et al. 1980).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences panicles.
subsp. chapmanii
1. Inflorescences racemes.
→ 2
2. Pistillate pedicels 0.5–3 cm; phyllodia less than 1 cm wide
subsp. graminea
2. Pistillate pedicels 2.1–5 cm; phyllodia more than 1 cm wide
subsp. weatherbiana
Source FNA vol. 22. FNA vol. 22.
Parent taxa Alismataceae > Sagittaria Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
S. ambigua, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
S. graminea subsp. chapmanii, S. graminea subsp. graminea, S. graminea subsp. weatherbiana
Synonyms S. longirostra var. australis, S. engelmanniana subsp. longirostra
Name authority Michaux: Flora Boreali-Americana 2: 190. (1803) (J. G. Smith) Small: Flora of the Southeastern United States 45. (1903)
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