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arum-leaf arrowhead, northern arrowhead, sagittaria cuneaire, tule potato, wapato

arrowhead family, water-plantain family

Habit Herbs, perennial, to 110 cm; rhizomes absent; stolons present; corms present. Herbs, annual or perennial, rhizomatous, stoloniferous, or cormose, caulescent, glabrous to stellate-pubescent; sap milky.
Roots

septate or not septate.

Leaves

emersed, floating, and submersed; submersed phyllodial, flattened, to 45 cm; floating with petiole triangular, to 100 cm, blade cordate or sagittate, rarely linear or ovate, 7.5–9 × 3.5–4 cm; emersed with petiole recurved, 3.5–51 cm, blade linear to sagittate, 2.5–17 × 1.5–11 cm, basal lobes when present shorter than remainder of blade.

basal, submersed, floating, or emersed, sessile or petiolate, sheathing proximally;

blade with translucent markings of dots or lines present or absent, basal lobes present or absent;

venation reticulate, primary veins parallel from base of blade to apex, secondary veins reticulate.

Inflorescences

racemes, rarely panicles, of 2–10 whorls, emersed, 14–21 × 2–10 cm, peduncle triangular, 10–50 cm;

bracts connate more than or equal to ¼ total length, lance-attenuate or acute, mostly (4–)7–40 mm, membranous, not papillose; fruiting pedicels ascending, cylindric, 0.5–2 cm.

scapose racemes or panicles, rarely umbels, erect, rarely floating or decumbent, whorled (forming racemes) or whorls branching (forming panicles), bracteolate.

Flowers

to 25 mm diam.;

sepals recurved, not enclosing flower or fruiting head;

filaments not dilated, equal to or longer than anthers, glabrous;

pistillate pedicellate, without ring of sterile stamens.

bisexual or unisexual, if unisexual, staminate and pistillate on same or different plants, hypogynous, subsessile to long-pedicellate;

sepals persistent, 3;

petals deciduous, 3, delicate;

stamens 0, 6, 9, or to 30, distinct;

anthers 2-loculed, dehiscing longitudinally;

pistils 0 or 6–1500 or more, distinct or coherent proximally, 1-loculed;

placentation basal;

ovules1–2.

Fruits

achenes or follicles.

Seeds

embryo U-shaped;

endosperm absent in mature seed.

Fruiting

heads 0.8–1.5 cm diam.;

achenes obovoid, abaxially keeled, 1.8–2.6 × 1.3–2.5 mm, beaked;

face not tuberculate, wings 0–1, ± entire, glands 0–1;

beak apical, erect, 0.1–0.4 mm.

2n

= 22.

Sagittaria cuneata

Alismataceae

Phenology Flowering late spring–summer (Jun–Sep).
Habitat Calcareous and muddy shores and shallow waters of rivers, lakes, ponds, pastures, and ditches, occasional in tidal waters, or in deep flowing water with slow current
Elevation 100–2500 m (300–8200 ft)
Distribution
from FNA
AK; AZ; CA; CO; CT; IA; ID; IL; IN; KS; MA; ME; MI; MN; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; VT; WA; WI; WY; AB; BC; LB; MB; NB; NS; NT; ON; QC; SK; YT
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Nearly worldwide; primarily tropical and subtropical regions
[BONAP county map]
Discussion

Sagittaria cuneata is extremely variable. On emersed plants, the leaf petioles are often bent toward the ground. Submersed plants often grow from a basal rosette with a long flexuous petiole and a floating sagittate leave. Plants in deep rivers often develop broad, straplike phyllodia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 12, species ca. 80 (4 genera, 34 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistils weakly coherent proximally into starlike aggregation; petals erose
Damasonium
1. Pistils distinct, forming heads or rings; petals entire.
→ 2
2. Pistils arranged in ring around margin of flattened receptacle.
Alisma
2. Pistils spirally arranged on convex receptacle.
→ 3
3. Flowers all bisexual; fruits mostly plump, longitudinally ribbed, lateral wings absent.
Echinodorus
3. Flowers unisexual (at least the proximal); fruits compressed, lateral wing often present, 1, curved.
Sagittaria
Source FNA vol. 22. FNA vol. 22, p. 7. Authors: Robert R. Haynes, C. Barre Hellquist.
Parent taxa Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. australis, S. brevirostra, S. cristata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
Alisma, Damasonium, Echinodorus, Sagittaria
Synonyms S. arifolia
Name authority E. Sheldon: Bulletin of the Torrey Botanical Club 20:283, plate 159. (1893) Ventenat
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