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Appalachian arrowhead, longbeak arrowhead

arrowhead family, water-plantain family

Habit Herbs, perennial, to 130 cm; rhizomes absent; stolons present; corms present. Herbs, annual or perennial, rhizomatous, stoloniferous, or cormose, caulescent, glabrous to stellate-pubescent; sap milky.
Roots

septate or not septate.

Leaves

emersed;

petiole 5-winged, 19–85 cm;

blade sagittate, 3–19 × 2.5–11 cm, basal lobes ± equal to remainder of blade.

basal, submersed, floating, or emersed, sessile or petiolate, sheathing proximally;

blade with translucent markings of dots or lines present or absent, basal lobes present or absent;

venation reticulate, primary veins parallel from base of blade to apex, secondary veins reticulate.

Inflorescences

racemes, of 5–12 whorls, emersed, 10–29 × 3–5 cm;

peduncles 25–105 cm;

bracts distinct or if connate, then less than ¼ total length, lanceolate, 7–30 mm, papery, not papillose; fruiting pedicels spreading to ascending, cylindric, 0.3–2.3 cm.

scapose racemes or panicles, rarely umbels, erect, rarely floating or decumbent, whorled (forming racemes) or whorls branching (forming panicles), bracteolate.

Flowers

to 3 cm diam.;

sepals recurved to spreading, not enclosing flower or fruiting head;

filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, without ring of sterile stamens.

bisexual or unisexual, if unisexual, staminate and pistillate on same or different plants, hypogynous, subsessile to long-pedicellate;

sepals persistent, 3;

petals deciduous, 3, delicate;

stamens 0, 6, 9, or to 30, distinct;

anthers 2-loculed, dehiscing longitudinally;

pistils 0 or 6–1500 or more, distinct or coherent proximally, 1-loculed;

placentation basal;

ovules1–2.

Fruits

achenes or follicles.

Seeds

embryo U-shaped;

endosperm absent in mature seed.

Fruiting

heads 1–2.2 cm diam.;

achenes obovoid, without abaxial keel, 2.1–3.2 × 1.4–2.3 mm, beaked;

faces not tuberculate, wings 0–2, ± entire, glands absent;

beak lateral, strongly recurved, 4–17 mm.

2n

= 22.

Sagittaria australis

Alismataceae

Phenology Flowering summer–early fall (Jul–Oct).
Habitat Slightly basic to slightly acidic ponds, lakes, and swamps
Elevation 1–300 m (0–1000 ft)
Distribution
from FNA
AL; AR; FL; GA; IA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Nearly worldwide; primarily tropical and subtropical regions
[BONAP county map]
Discussion

The name Sagittaria longirostra (Micheli) J. G. Smith has been misapplied to S. australis (J. G. Smith) Small (E. O. Beal et al. 1980).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 12, species ca. 80 (4 genera, 34 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistils weakly coherent proximally into starlike aggregation; petals erose
Damasonium
1. Pistils distinct, forming heads or rings; petals entire.
→ 2
2. Pistils arranged in ring around margin of flattened receptacle.
Alisma
2. Pistils spirally arranged on convex receptacle.
→ 3
3. Flowers all bisexual; fruits mostly plump, longitudinally ribbed, lateral wings absent.
Echinodorus
3. Flowers unisexual (at least the proximal); fruits compressed, lateral wing often present, 1, curved.
Sagittaria
Source FNA vol. 22. FNA vol. 22, p. 7. Authors: Robert R. Haynes, C. Barre Hellquist.
Parent taxa Alismataceae > Sagittaria
Sibling taxa
S. ambigua, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres
Subordinate taxa
Alisma, Damasonium, Echinodorus, Sagittaria
Synonyms S. longirostra var. australis, S. engelmanniana subsp. longirostra
Name authority (J. G. Smith) Small: Flora of the Southeastern United States 45. (1903) Ventenat
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