arrowhead, sagittaire
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annual sagittaria, California arrowhead, California lophotocarpus, giant arrowhead, hooded arrowhead, sagittaire spongieuse, spongy-leaf arrowhead
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Plants perennial, rarely annual, submersed, floating-leaved, or emersed, glabrous to sparsely pubescent; rhizomes often present, occasionally terminated by tubers; stolons often present; corms absent; tubers white to brown, smooth. |
Herbs, annual or perennial, to 100 cm; rhizomes present; stolons present; corms present. |
septate. |
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sessile or petiolate; petiole terete to triangular; blade with translucent markings absent, linear to obovate, base attenuate to hastate or sagittate, margins entire, apex round to acute. |
submersed and emersed; submersed leaves absent or phyllodial, flattened, to 17 × 2 cm; emersed leaves with petiole triangular, 21–55 cm, blade hastate to sagittate, 2.5–17.5 × 0.6–22 cm, lobes longer than or equal to remainder of blade. |
racemes, panicles, rarely umbels, of 1–17 whorls, erect, emersed or floating, rarely submersed; bracts coarse or delicate, apex obtuse to acute, smooth or papillose proximally to distally. |
racemes or panicles, of 1–15 whorls, floating or emersed, 1.5–28 × 1.5–15 cm; peduncles 15–47 cm; bracts distinct or connate less than ¼ total length, lanceolate to elliptic, 4–34 mm, delicate, not papillose; fruiting pedicels recurved, club-shaped, 0.5–4.2 cm. |
unisexual, the proximal rarely with ring of sterile stamens; staminate flowers pedicellate, distal to pistillate flowers; pistillate flowers mostly pedicellate, rarely sessile; bracts subtending pedicels, lanceolate, shorter than pedicels, apex obtuse to acute; pedicels ascending to recurved; receptacle convex; sepals recurved in staminate flowers, recurved to erect in pistillate flowers, often sculptured, herbaceous to leathery; petals white, rarely with pink spot or tinge, entire; stamens 7–30; filaments linear to dilated, glabrous to pubescent; pistils to 1500 or more, spirally arranged, not radiating in starlike pattern, distinct; ovules 1; styles terminal. |
2–5 cm diam.; sepals spreading in staminate, erect in pistillate, enclosing flower or fruiting head; filaments cylindric, longer than anthers, glabrous; pistillate pedicellate, with or without ring of sterile stamens. |
without longitudinal ribs, compressed, abaxially keeled or not, abaxial wings often present, lateral wing often present, 1, curved, glands present. |
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heads 1.2–2.1 cm diam.; achenes oblanceoloid, not abaxially keeled, 2–4.3 × 0.7–1.5 mm, beaked; faces not tuberculate, wings absent, glands 1; beak lateral, horizontal, 0.4–0.8 mm. |
= 11. |
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Mostly Western Hemisphere; Europe; Asia |
AL; AR; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; SC; SD; TN; TX; VA; WI; WV; NB; Mexico; South America
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Species ca. 30 (24 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Fruiting pedicels recurved or rarely spreading; pistillate sepals mostly erect and closely enclosing flower or fruiting head, occasionally spreading to recurved. | → 2 |
1. Fruiting pedicels spreading to ascending or absent; pistillate sepals mostly spreading to recurved, not enclosing flower. | → 9 |
2. Leaves, at least some, emersed. | → 3 |
2. Leaves submersed, floating, rarely emersed, or plants stranded on shore. | → 5 |
3. Fruiting heads 1.2–2.1 cm diam.; leaf blade hastate to sagittate | S. montevidensis |
3. Fruiting heads 0.7–1.2 cm diam.; leaf blade linear-ovate to lance-elliptic. | → 4 |
4. Achenes abaxially keeled, mostly more than 2 mm; faces not tuberculate, often with glands; California. | S. sanfordii |
4. Achenes not abaxially keeled, mostly less than 2 mm, faces tuberculate, without glands; e of Rocky Mountains. | S. platyphylla |
5. Achene faces tuberculate; floating leaves with sagittate blades present at least on some plants of population. | S. guayanensis |
5. Achene faces not tuberculate; floating leaves absent or, if present, with unlobed or hastate blades. | → 6 |
6. Fruiting pedicels 0.2–1.1 cm; phyllodia lenticular in cross section; tidal muds; mostly brackish waters. | S. subulata |
6. Fruiting pedicels 1.5–6.5 cm; phyllodia flattened in cross section; rarely tidal muds; mostly fresh waters. | → 7 |
7. Leaves with blades and petioles usually present on some plants; some plants often stranded along shore, these usually with expanded leaf blades. | S. filiformis |
7. Leaves all phyllodia; plants almost always submersed, rarely stranded, these without expanded leaf blades. | → 8 |
8. Phyllodia 50–250 cm; Florida springs. | S. kurziana |
8. Phyllodia 12–53 cm; streams and lakes, New Mexico. | S. demersa |
9. Filaments pubescent to tomentulose (except S. fasciculata). | → 10 |
| → 17 |
10. Leaves all phyllodia, nearly terete; ne United States. | S. teres |
10. Leaves with blades and petioles or if phyllodia, then flattened to triangular in cross section; mostly widespread. | → 11 |
11. Pistillate flowers sessile to subsessile. | S. rigida |
11. Pistillate flowers obviously pedicellate. | → 12 |
| S. lancifolia |
| → 13 |
13. Rhizomes present, coarse; stolons and corms absent. | → 14 |
13. Rhizomes absent or if present, then not coarse; corms and/or stolons present. | → 15 |
14. Abaxial wing of fruit ± entire; plants widespread | S. graminea |
14. Abaxial wing of fruit scalloped or toothed; ne Alabama | S. secundifolia |
15. Filaments exceeding anthers in length. | S. cristata |
15. Filaments shorter than or equaling anthers in length. | → 16 |
16. Blades of emersed leaves 0.5 cm or more wide; w Carolinas. | S. fasciculata |
16. Blades of emersed leaves, if present, 0.4(–0.5) cm or less wide; se coastal plain. | S. isoetiformis |
17. Emersed leaf blades linear to ovate. | → 18 |
17. Emersed leaf blades cordate, sagittate, or hastate. | → 20 |
| S. papillosa |
18. Bracts not papillose. | → 19 |
19. Emersed plants with erect to ascending petioles; leaf blades lanceolate to ovate. | S. ambigua |
19. Emersed plants with recurved petioles; leaf blades linear to sagittate. | S. cuneata |
20. Bracts distinct or connate much less than ¼ total length. | → 21 |
20. Bracts connate at least ¼ total length. | → 23 |
21. Flowers in 2–4 whorls; achenes with facial glands. | S. engelmanniana |
21. Flowers in 5–12 whorls; achenes without facial glands. | → 22 |
22. Petiole winged in cross section; achene beak strongly recurved. | S. australis |
22. Petiole ridged in cross section; achene beak ascending apically. | S. brevirostra |
23. Achene beak 1–2 mm, horizontal. | S. latifolia |
23. Achene beak 0.1–0.6 mm, erect or incurved. | → 24 |
24. Emersed plants with recurved petioles and linear to sagittate blades; basal lobes equal to or shorter than remainder of blade; submersed leaves phyllodial, floating leaves cordate to sagittate | S. cuneata |
24. Emersed plants with ascending to erect petioles and sagittate blades; basal lobes longer than remainder of blade; submersed and floating leaves absent. | S. longiloba |
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1. Plants of tidal mud flats, often stranded by low tide, emersed blades spatulate, rarely sagittate. | subsp. spongiosa |
1. Plants of fresh waters, not stranded by low tide; emersed blades hastate to sagittate. | → 2 |
2. Pistillate flowers without ring of sterile stamens; petals with purple spot at base | subsp. montevidensis |
2. Pistillate flowers with ring of sterile stamens; petals without purple spot at base. | subsp. calycina |
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FNA vol. 22. |
FNA vol. 22. |
Alismataceae |
Alismataceae > Sagittaria |
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S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres |
S. ambigua, S. australis, S. brevirostra, S. cristata, S. cuneata, S. demersa, S. engelmanniana, S. fasciculata, S. filiformis, S. graminea, S. guayanensis, S. isoetiformis, S. kurziana, S. lancifolia, S. latifolia, S. longiloba, S. montevidensis, S. papillosa, S. platyphylla, S. rigida, S. sanfordii, S. secundifolia, S. subulata, S. teres |
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Lophotocarpus |
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Linnaeus: Sp. Pl. 2: 993. 1753; Gen. Pl. ed. 5; 429, (1754) |
Chamisso & Schlechtendal: Linnaea 2: 156. (1827) |
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