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coastal pearlwort, fleshy pearlwort, stick-stem pearlwort, sticky-stem pearlwort, thick-stem pearlwort

alpine pearlwort, arctic pearlwort, sagine des alpes

Habit Plants annual or perennial, tufted, glabrous or glandular-pubescent. Plants perennial, tufted or becoming cespitose in alpine habitats, glabrous.
Stems

spreading to decumbent or procumbent, much-branched, stout, rarely filiform, distal portion glandular-pubescent.

ascending or sometimes procumbent, few- to many-branched, not filiform.

Leaves

axillary fascicles absent;

basal rosette or tuft of ascending leaves usually present;

basal blades linear, 10–30 mm, succulent, apex apiculate, glabrous;

cauline leaves conspicuously connate basally, forming shallow, scarious cup, blade linear, fleshy, apex apiculate, glabrous;

proximal blades 6–15(–20) mm, distal blades rarely subulate, (2.5–)3.5–7(–9) mm.

axillary fascicles absent;

basal frequently in primary and secondary rosettes 9–45 mm diam., blade linear, 10–20 mm, not succulent, apex apiculate, rarely aristate, glabrous;

cauline not conspicuously connate basally, rarely forming inflated cup in cespitose, alpine plants, blade linear, sometimes linear-subulate in cespitose plants, 4–20(–25) mm, not fleshy, apex apiculate, glabrous.

Pedicels

slender to stout, glabrous or glandular-pubescent distally.

frequently recurved during capsular development, erect in fruit, filiform, glabrous.

Flowers

axillary, 5-merous;

calyx bases glabrous or glandular-pubescent;

sepals ovate to orbiculate, (2–)2.5–3.5 mm, hyaline margins whitish, occasionally purple tinged on margins or apex, apex obtuse to rounded, glabrous or glandular-pubescent, remaining appressed following capsule dehiscence;

petals elliptic to nearly orbiculate, (1.5–)2–2.5(–3) mm, shorter than sepals;

stamens 10.

axillary or terminal, 5-merous, very rarely some 4-merous;

calyx base glabrous;

sepals elliptic, 2–2.5 mm, hyaline margins white, rarely purple in alpine specimens, apex obtuse to rounded, remaining appressed following capsule dehiscence;

petals elliptic, (1–)1.5–2 mm, shorter than or equaling sepals;

stamens (5 or) 10.

Capsules

(3–)3.5–4.5 mm, exceeding sepals, dehiscing ca. 1/4 length.

2.5–3(–3.5) mm, 1.5–2 times sepals, dehiscing to base.

Seeds

reddish brown, reniform with abaxial groove absent, plump, 0.5 mm, smooth or slightly pebbled.

brown, obliquely triangular with distinct abaxial groove, 0.3–0.4 mm, smooth to slightly pebbled.

2n

= 22.

Sagina maxima

Sagina saginoides

Phenology Flowering mid-late summer.
Habitat Montane sites, open or light shade, wet places on lake margins, along stream gravels and seepages in rock ledges and roadcuts, subalpine and alpine zones
Elevation 1000-4000 m (3300-13100 ft)
Distribution
from FNA
AK; CA; OR; WA; BC; e Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; NF; NT; NU; QC; YT; Mexico; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some specimens from alpine habitats in Montana and Alberta are intermediate between Sagina saginoides and the typically arctic S. nivalis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx bases and distal portion of stems and pedicels glandular-pubescent
subsp. maxima
1. Calyx bases, stems, and distal portion of pedicels entirely glabrous
subsp. crassicaulis
Source FNA vol. 5, p. 146. FNA vol. 5, p. 142.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Sagina Caryophyllaceae > subfam. Alsinoideae > Sagina
Sibling taxa
S. apetala, S. caespitosa, S. decumbens, S. japonica, S. nivalis, S. nodosa, S. procumbens, S. saginoides, S. subulata
S. apetala, S. caespitosa, S. decumbens, S. japonica, S. maxima, S. nivalis, S. nodosa, S. procumbens, S. subulata
Subordinate taxa
S. maxima subsp. crassicaulis, S. maxima subsp. maxima
Synonyms Spergula saginoides, S. linnaei, S. micrantha, S. saginoides var. hesperia
Name authority A. Gray: Mem. Amer. Acad. Arts, n. s. 6: 382. (1858) (Linnaeus) H. Karsten: Deut. Fl., 539. (1882)
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