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coastal pearlwort, fleshy pearlwort, stick-stem pearlwort, sticky-stem pearlwort, thick-stem pearlwort

Photo is of parent taxon

beach pearlwort, coastal pearlwort, fleshy pearlwort, stick-stem pearlwort, stickystem pearlwort, thick-stem pearlwort

Habit Plants annual or perennial, tufted, glabrous or glandular-pubescent. Plants perennial, glabrous or mostly so.
Stems

spreading to decumbent or procumbent, much-branched, stout, rarely filiform, distal portion glandular-pubescent.

spreading, decumbent, or procumbent, glabrous, nodes frequently purple tinged.

Leaves

axillary fascicles absent;

basal rosette or tuft of ascending leaves usually present;

basal blades linear, 10–30 mm, succulent, apex apiculate, glabrous;

cauline leaves conspicuously connate basally, forming shallow, scarious cup, blade linear, fleshy, apex apiculate, glabrous;

proximal blades 6–15(–20) mm, distal blades rarely subulate, (2.5–)3.5–7(–9) mm.

basal leaves in rosette of broadly linear, fleshy leaves, or absent with primary or secondary tufts of ascending, linear basal leaves, these usually less fleshy than rosette leaves (rosettes rarely present in plants occurring north of Washington);

cauline leaf blades: proximal 6–15 mm, distal 3–5 mm, glabrous.

Pedicels

slender to stout, glabrous or glandular-pubescent distally.

slender to stout, glabrous.

Flowers

axillary, 5-merous;

calyx bases glabrous or glandular-pubescent;

sepals ovate to orbiculate, (2–)2.5–3.5 mm, hyaline margins whitish, occasionally purple tinged on margins or apex, apex obtuse to rounded, glabrous or glandular-pubescent, remaining appressed following capsule dehiscence;

petals elliptic to nearly orbiculate, (1.5–)2–2.5(–3) mm, shorter than sepals;

stamens 10.

calyx glabrous;

sepals ovate to nearly orbiculate, (2–)2.5–3(–3.5) mm;

petals elliptic to orbiculate, (1.5–)2–2.5(–3) mm, slightly shorter than sepals.

Capsules

(3–)3.5–4.5 mm, exceeding sepals, dehiscing ca. 1/4 length.

(3–)3.5–4(–4.5) mm.

Seeds

reddish brown, reniform with abaxial groove absent, plump, 0.5 mm, smooth or slightly pebbled.

smooth to slightly pebbled.

2n

= 46, 66.

Sagina maxima

Sagina maxima subsp. crassicaulis

Phenology Flowering spring–early autumn.
Habitat Coastal, moist, sandy bluffs, crevices of rock cliffs, at or near high-tide mark, gravelly-sandy beaches
Elevation 0-10 m (0-0 ft)
Distribution
from FNA
AK; CA; OR; WA; BC; e Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; OR; WA; BC; Asia (Kamchatka)
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Integradation occurs where the range of subsp. crassicaulis overlaps with that of subsp. maxima. Variation of pubescence in populations on Vancouver Island and the Queen Charlotte Islands ranges from completely glabrous specimens typical of subsp. crassicaulis to individuals with pedicels and calyx bases weakly pubescent, to others with densely pubescent pedicels. Subspecies crassicaulis is far more common than subsp. maxima.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx bases and distal portion of stems and pedicels glandular-pubescent
subsp. maxima
1. Calyx bases, stems, and distal portion of pedicels entirely glabrous
subsp. crassicaulis
Source FNA vol. 5, p. 146. FNA vol. 5, p. 147.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Sagina Caryophyllaceae > subfam. Alsinoideae > Sagina > Sagina maxima
Sibling taxa
S. apetala, S. caespitosa, S. decumbens, S. japonica, S. nivalis, S. nodosa, S. procumbens, S. saginoides, S. subulata
S. maxima subsp. maxima
Subordinate taxa
S. maxima subsp. crassicaulis, S. maxima subsp. maxima
Synonyms S. crassicaulis
Name authority A. Gray: Mem. Amer. Acad. Arts, n. s. 6: 382. (1858) (S. Watson) G. E. Crow: Rhodora 80: 79. (1978)
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