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ditch grass, spiral ditch-grass

ditch-grass, rruppia, widgeon grass, widgeonweed

Habit Herbs, rooting at proximal nodes.
Stems

to 55 cm × 0.1–0.3 mm.

Leaves

3.2–45.1 cm;

blade 0.2–0.5 mm wide, apex acute.

blade entire proximally, minutely serrulate distally, apex ± obtuse to acute;

veins 1.

Inflorescences

peduncles with 5–30 coils, 30–300 × 0.5 mm.

fewer than 20-flowered, at first enclosed by sheathing leaf bases.

Flowers

pistils 4–6.

bisexual;

anthers 2-loculed, locules separated by broad connective;

stipe elongating after anthesis.

Fruits

1.5–2 × 1.1–1.5 mm;

gynophore 2–3.5 cm;

beak lateral, erect, 0.5–1 mm.

beaked, long-stipitate;

beak erect or slightly recurved.

2n

= 40 (Europe).

Ruppia cirrhosa

Ruppia

Phenology Flowering summer–fall.
Habitat Shallow to deep fresh waters of lakes with high concentrations of sulfur or calcium
Elevation 300–2500 m (1000–8200 ft)
Distribution
from FNA
AK; AZ; CA; CO; IL; KS; MI; MN; MO; MT; ND; NE; NM; NV; OH; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; NT; SK; YT; Central America; South America; West Indies; Europe
[WildflowerSearch map]
[BONAP county map]
from USDA
Almost worldwide
[BONAP county map]
Discussion

Ruppia spiralis Linnaeus has occasionally been used for this taxon in North America. Linnaeus, however, had never published that name (J. T. A. Verhoeven 1979). Ruppia cirrhosa is, indeed, the correct name for the taxon (J. C. Gamerro 1968).

An implication that the North American material with long, spiraling peduncles is different from the European material of Ruppia cirrhosa stems from the statement that R. occidentalis occurs in inland lakes (J. T. A. Verhoeven 1979). No differences between European R. cirrhosa and the North American material are listed, however. In fact, a comparison of the Verhoeven's descriptive discussions for of R. cirrhosa (Verhoeven 1979) with the North American material shows that the two are the same. The genus should be studied on a worldwide basis. Until further studies indicate otherwise, I am considering North American and European material to be conspecific.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Considerable confusion exists about North American taxa of Ruppia. Two distinct forms, along with several intermediates, are known: one with short peduncles with four or fewer spirals and another with long peduncles with five or more coils (often many more). Usually, those with few coils are in brackish waters near the coast, whereas those with many coils are inland, often in lakes that have high mineral contents. The forms have been considered variants variants of one species, R. maritima (M. L. Fernald and K. M. Wiegand 1914); more recently they have been accepted at species level (R. F. Thorne 1993). I am adopting Thorne’s concepts.

Species ca. 10 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Peduncle with 5 or more coils in fruit, longer than 30 mm; plants mostly from inland localities
R. cirrhosa
1. Peduncle with fewer than 5 coils in fruit, shorter than 25 mm; plants mostly from coastal localities
R. maritima
Source FNA vol. 22. FNA vol. 22.
Parent taxa Ruppiaceae > Ruppia Ruppiaceae
Sibling taxa
R. maritima
Subordinate taxa
R. cirrhosa, R. maritima
Synonyms Buccaferrea cirrhosa, R. cirrhosa subsp. occidentalis, R. occidentalis
Name authority (Petagna) Grande: Bull. Orto Bot. Regia Univ. Naapoli 5: 58. (1918) Linnaeus: Sp. Pl. 1: 127. 1753;, Gen. Pl. ed. 5; 61, (1754)
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