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Rumex venosus

veiny dock, wild begonia, wing dock

arctic dock

Habit Plants perennial, glabrous or nearly so, with creeping rhizomes. Plants perennial, glabrous or nearly so, with fusiform, oblique rootstock, occasionally with horizontal, short-creeping rhizome.
Stems

ascending or, rarely, erect, usually producing axillary shoots near base, (10–)15–30(–40) cm.

erect, simple or branched in distal 2/3 (then with few, comparatively short branches), 10–70(–100) cm.

Leaves

blades ovate-elliptic, obovate-elliptic, or ovate-lanceolate, (2–)4–12(–15) × 1–5(–6) cm, subcoriaceous, base narrowly to broadly cuneate, margins entire, flat or slightly undulate, apex acute or acuminate.

ocrea deciduous or partially persistent at maturity;

blade narrowly lanceolate, lanceolate, or oblong-lanceolate, normally 5–15(–20) × 1.5–5 cm, base cuneate to broadly cuneate, rarely truncate or very weakly cordate, margins entire or rarely indistinctly repand, flat, apex acute or subacute.

Inflorescences

terminal and axillary, usually occupying distal 2/3 of stem/shoot, usually dense, or interrupted in proximal part, broadly paniculate.

terminal, occupying distal 1/2–2/3 of stem, interrupted, paniculate, simple or nearly so (branches, when present usually less than 7–8 cm).

Pedicels

articulated near middle, filiform or slightly thickened, (8–)10–16 mm, articulation distinct, slightly swollen.

articulated in proximal 1/3, filiform, 5–13(–17) mm, usually not more than 2–2.5 times as long as inner tepals, articulation weakly evident, not swollen.

Flowers

5–15 in whorls;

inner tepals distinctly double-reticulately veined, orbiculate or reniform-orbiculate, 13–18(–20) × (20–)23–30 mm, base deeply emarginate or cordate, margins entire, apex rounded, obtuse, rarely subacute, with short, broadly triangular tip;

tubercles absent, occasionally very small.

7–15 in whorls;

inner tepals ovate, 4.5–7.5(–8) × 4–6(–7) mm, base truncate to weakly cordate, margins entire, apex obtuse or subacute;

tubercles absent.

Achenes

brown or dark brown, 5–7 × 4–6 mm.

reddish brown, 3–4 × 1.5–2 mm.

2n

= 40.

= 40, 120, ca. 170, ca. 200.

Rumex venosus

Rumex arcticus

Phenology Flowering spring–early summer. Flowering late spring–summer.
Habitat Sand dunes, sandy and gravelly riverbanks and slopes, deserts, grasslands 200-1500 m Moist tundra, marshes, river valleys, sandy and gravelly shores and slopes
Elevation 0-1000 m [0-3300 ft]
Distribution
from FNA
AZ; CA; CO; IA; ID; KS; MN; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WI; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; NT; NU; YT; ne Europe; n Asia (arctic and subarctic zones)
[WildflowerSearch map]
[BONAP county map]
Discussion

Rumex venosus is a distinctive species rarely confused with any other members of the genus. However, I have seen herbarium specimens of it misidentified as R. hymenosepalus, and vice versa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rumex arcticus is polymorphic, as are R. aquaticus in the narrow sense and R. occidentalis. However, unlike R. aquaticus and R. occidentalus, it is represented by at least two chromosome races (G. A. Mulligan and C. Frankton 1972; Á. Löve 1986). Plants morphologically transitional between R. arcticus and R. aquaticus were described from Kamchatka as R. kamtschadalus (= R. arcticus var. kamtschadalus). The same forms occasionally occur in northwestern North America. According to Á. Löve and D. Löve (1975b) and Á. Löve (1986), they are usually tetraploids (2n = 40) and deserve recognition at the species level. However, they are not always morphologically distinct from R. arcticus. The group needs additional study, and at present I prefer to keep the tetraploid plants provisionally within R. arcticus, regarding them as var. kamtschadalus.

A few highly sterile specimens with mostly abortive flowers, which I have seen in Alaskan herbarium material, most probably represent hybrids between tetraploid and 12-ploid races of the R. aquaticus group.

Plants with unusually wide, triangular-oblong, or almost ovate leaves were described as var. latifolius Tolmatchew. This seems to be a predominant variety on the Beringian coast of Chukotka and Wrangel and Ratmanov islands (A. I. Tolmachew 1966). I also have seen at least two collections of this variety from the western coast of Alaska. The enigmatic var. perlatus Hultén may belong here (see discussion under 8. R. lapponicus).

Rumex arcticus has been reported from Churchill in northeastern Manitoba (H. J. Scoggan 1978–1979, part 3). That record needs confirmation because some northern forms of R. arcticus and R. occidentalis are similar.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Axillares Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Rumex
Sibling taxa
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. arcticus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hastatulus, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. maritimus, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. verticillatus, R. violascens
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hastatulus, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. maritimus, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. venosus, R. verticillatus, R. violascens
Synonyms R. aquaticus subsp. arcticus, R. arcticus var. kamtschadalus, R. arcticus var. latifolius, R. domesticus var. nanus, R. kamtschadalus, R. longifolius var. nanus, R. ursinus
Name authority Pursh: Fl. Amer. Sept. 2: 733. (1813) Trautvetter: in A. T. von Middendorff, Reise Siber. 1(2,1): 29. (1847)
Source FNA vol. 5, p. 505. Treatment author: Sergei L. Mosyakin. FNA vol. 5, p. 518. Treatment author: Sergei L. Mosyakin.
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