Rumex spiralis |
Rumex lapponicus |
|
---|---|---|
spiral tall dock, wing dock |
dock, green sorrel, Lapland dock, Lapland Mountain sorrel, Lapland or Lapland Mountain sorrel |
|
Habit | Plants perennial, glabrous, with creeping rhizomes. | Plants perennial, glabrous or nearly so, with short and relatively thin, horizontal or slightly oblique rootstock (usually not reaching deep into substrate) and ± crowded 2d-order roots (occasionally with short-creeping rhizome). |
Stems | ascending or erect, usually producing axillary shoots below 1st-order inflorescence or at proximal nodes, 50–90 cm. |
erect, rarely ascending, several from base or often solitary, branched in distal 1/2 (in inflorescence), (10–)20–60(–100) cm. |
Leaves | blades ovate-lanceolate, oblong-lanceolate, or lanceolate, 10–15 × 3–5.5 cm, usually 2.5–3.5 times as long as wide, widest in proximal 1/3, thick, usually not coriaceous, base broadly cuneate, truncate, or rounded, margins entire, flat or slightly undulate-crisped, apex acute or attenuate. |
ocrea of at least middle and distal leaves with margins entire, not fringed, sometimes laciniate but only in distal parts; blade broadly ovate, rarely almost round, oblong-ovate, rarely oblong-lanceolate, 3–10(–14) × 1–4(–5) cm, normally less than 2.5 times as long as wide, base sagittate (with acute or subacute lobes directed downward, ± parallel to petiole, or slightly incurved inward), margins entire, normally flat, apex subacute or obtuse. |
Inflorescences | terminal and axillary, terminal usually occupying distal 1/2 of stem, dense, narrowly to broadly paniculate (branches usually simple). |
terminal, occupying distal 1/3 of stem, usually lax and interrupted especially in proximal part, narrowly paniculate or occasionally simple, cylindric (with 1st-order branches simple, or with few 2d-order branches). |
Pedicels | articulated in proximal 1/3, thin but slightly thickened distally, (2–)3–7(–8) mm, usually as long as or shorter than inner tepals, articulation slightly swollen. |
articulated near middle, filiform, 2–5 mm, articulation distinct. |
Flowers | 12–20 in whorls; inner tepals broadly cordate or broadly ovate-deltoid, 7–10 × 8–12 mm, base deeply and broadly cordate, margins entire, apex acuminate; tubercles 3, equal or subequal, usually minutely to distinctly rugose. |
(2–)4–8 in whorls; inner tepals orbiculate, occasionally broadly ovate, 3.5–4.5 × 3.5–4.5 mm, base rounded or cordate, apex obtuse; tubercles small or occasionally absent. |
Achenes | brown or dark reddish brown, 2.5–3.5 × 2–2.5 mm. |
brown or dark brown to brownish yellow, 1.7–2.5 × 0.9–1.3 mm, dull. |
2n | = 20. |
= 14 (pistillate plants), 15 (staminate plants). |
Rumex spiralis |
Rumex lapponicus |
|
Phenology | Flowering spring. | Flowering late spring–summer. |
Habitat | Sandy and gravelly shores | Meadows, rock outcrops, alluvial habitats along rivers and streams in tundra and montane and subalpine zones |
Elevation | 0-200 m (0-700 ft) | 0-2500 m (0-8200 ft) |
Distribution |
TX |
AK; MT; WY; AB; BC; NT; YT; Greenland; n Eurasia |
Discussion | Rumex spiralis is related to R. altissimus; however, it is geographically restricted and morphologically distinct. It has inner tepals larger than those of any other member of subsect. Salicifolii and distinctly wider leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Rumex acetosa has been commonly misapplied to R. lapponicus. Native North American montane plants of the R. acetosa aggregate from the Rocky Mountains southward to Beartooth Plateau in Montana and Wyoming usually were referred to as R. alpestris [= Acetosa pratensis Miller subsp. alpestris (Jacquin) Á. Löve; A. pratensis subsp. arifolia (A. Blytt & O. C. Dahl) Á. Löve; R. arifolius Allioni, not Linnaeus f.; R. acetosa subsp. alpestris (Jacquin) Á. Löve; R. acetosa subsp. arifolius A. Blytt & O. C. Dahl]. Recent nomenclatural studies demonstrated that R. alpestris is an ambiguous name, which was probably based on plants belonging to R. scutatus Linnaeus (see I. O. Pestova 1998), and accepted the name R. arifolius for the predominantly European montane taxon. It differs from arctic plants, as well as from montane forms of the R. acetosa aggregate from southern Siberia and temperate North America, by its more robust habit, more branched inflorescence (similar to that of R. thyrsiflorus), and larger and more acute triangular-sagittate leaves (see A. I. Tolmachew 1966; N. N. Tzvelev 1989b; Pestova; R. Elven et al. 2000). Montane races possibly developed independently from R. lapponicus-like or R. acetosa-like ancestors, and they are still unclear taxonomically. Because of that, I prefer to keep those forms within R. lapponicus. The whole aggregate needs careful study; however, some authors prefer to include all arcto-montane Holarctic races of this aggregate in the collective and rather polymorphic R. alpestris in the broad sense (see Á. Löve 1944; Löve and D. Löve 1957). Some arctic plants from western Alaska may be conspecific with Rumex pseudoxyria (Tolmatchew) Khokhrjakov [= R. acetosa subsp. pseudoxyria Tolmatchew; Acetosa pseudoxyria (Tolmatchew) Tzvelev], a taxon described from arctic eastern Siberia (A. I. Tolmachew 1966). This entity is evidently closely related to R. lapponicus but differs from all other members of the R. acetosa group in having basal leaves less than two times as long as wide, almost hastate or at least rounded-truncate at base, resembling those of Oxyria digyna (Linnaeus) Hill, cauline leaves small or completely reduced, and inflorescence occupying more than one-half of the stem; the plant itself is also somewhat similar in appearance to the European alpine species R. nivalis Hegetschweiler. According to E. Hultén (1973), his Rumex arcticus var. perlatus, described from a single specimen collected at Tin City, Seward Peninsula, Alaska, agrees perfectly with the original description of R. acetosa subsp. pseudoxyria. It is unlikely that Hultén would confuse two rather distantly related groups. There is also a possibility that var. perlatus is identical with R. arcticus var. latifolius Tolmatchew (see discussion under R. arcticus). Additional collections are needed to confirm the occurrence of R. pseudoxyria in northwestern North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 507. | FNA vol. 5, p. 503. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | R. acetosa subsp. lapponicus, Acetosa alpestris subsp. lapponica, Acetosa lapponica, R. alpestris subsp. lapponicus | |
Name authority | Small: Bull. Torrey Bot. Club 22: 44, plate 228. (1895) | (Hiitonen) Czernov: in B. N. Gorodkov and A. I. Pojarkova, Fl. Murmansk. Obl. 3: 154. (1956) |
Web links |