Rumex obtusifolius |
Polygonaceae subfam. polygonoideae |
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bitter dock, broad-leaf dock, broad-leaf or broadleaf or bitter dock, patience a feuilles obtuses |
knotweed |
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Habit | Plants perennial, glabrous or ± papillose especially on veins of leaf blades abaxially, with fusiform, vertical rootstock. | Trees, shrubs, vines, or herbs, perennial or annual, homophyllous (heretophyllous in some species of Polygonum); root fibrous or a solid taproot, rarely tuberous. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, branched distal to middle or occasionally in distal 2/3, often with few flowering stems from rootstock, 60–120(–150) cm. |
usually prostrate to erect, sometimes scandent, not scapose, rarely with recurved spines (some species of Persicaria), glabrous or pubescent, sometimes glandular; nodes usually swollen; branches free (adnate to stems distal to nodes and appearing to arise internodally in Polygonella); tendrils absent (except in Antigonon and Brunnichia). |
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Leaves | ocrea deciduous to partially persistent at maturity; blade oblong to ovate-oblong, sometimes broadly ovate, 20–40 × 10–15 cm, usually less than 4 times as long as wide, base normally distinctly cordate, occasionally rounded, rarely truncate, margins normally entire, flat or undulate, rarely slightly crisped, apex obtuse or subacute. |
deciduous (persistent in Coccoloba and sometimes more than 1 year in Antigonon and Polygonella), basal or basal and cauline, rarely cauline only, mostly alternate; ocrea present, persistent or deciduous, cylindric to funnelform, chartaceous, membranous, coriaceous, or, rarely, foliaceous or partly so; petiole present or absent, rarely articulate basally (Fagopyrum, Fallopia, Polygonella, Polygonum), rarely with extrafloral nectaries (Fallopia, Muehlenbeckia); blade simple with entire margins, rarely undulate or lobed. |
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Inflorescences | terminal, occupying distal 2/3 of stem, usually lax and interrupted, narrowly or broadly paniculate, branches usually forming angle of 30–45° with 1st-order stem. |
terminal or terminal and axillary, spikelike, racemelike, paniclelike, cymelike, or, rarely, capitate, comprising simple or branched clusters of compound inflorescences; bracts absent; peduncle spreading to erect, sometimes absent; clusters of flowers subtended by connate bracteoles forming persistent membranous tube (ocreola), awnless. |
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Pedicels | articulated in proximal 1/3 or rarely near middle, filiform, 2.5–8.5(–10) mm, articulation distinctly swollen. |
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Flowers | 10–25 in lax whorls; inner tepals ovate-triangular, deltoid or, occasionally, lingulate, 3–6 × 2–3.5 mm (excluding teeth), ca. 1.5–2 times as long as wide, base truncate, margins usually distinctly dentate, rarely subentire, apex obtuse to subacute, straight, teeth 2–5, normally at each side of margin, short-subulate or triangular-subulate, straight, 0.5–1.8 mm, or shorter than width of inner tepals; tubercle usually 1, sometimes 3, then 1 distinctly larger, smooth. |
usually bisexual, sometimes bisexual and unisexual on same plant, rarely unisexual only, 1–20+ per ocreate fascicle, often with stipelike base distal to articulation; perianth often accrescent in fruit, often greenish, white, pink, yellow, red, or purple, usually unwinged and unkeeled (winged or, sometimes, keeled in Fallopia, rarely keeled in Polygonum), campanulate or urceolate, sometimes membranous, indurate, or fleshy in fruit, rarely developing raised tubercles proximally (Rumex), glabrous or pubescent, sometimes glandular or glandular-punctate; tepals 2–6, usually in 2 whorls, distinct or connate proximally and forming tube, petaloid or sepaloid, monomorphic or dimorphic; nectary a disk at base of ovary or glands associated with bases of filaments; stamens usually (1–)6–9, staminodes rarely present; filaments distinct, or connate basally and sometimes forming staminal tube, free or adnate to perianth tube; pistils (2–)3(–4)-carpellate; ovary 1-locular (sometimes with vestigial partitions proximally); ovule 1, orthotropous or, rarely, anatropous, placentation basal or free-central; styles 1–3, erect to spreading or recurved, distinct or connate proximally; stigmas peltate, capitate, fimbriate, or penicillate. |
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Achenes | brown to reddish brown, 2–2.7 × 1.2–1.7 mm. |
yellowish, brown, red, or black, homocarpic (sometimes heterocarpic in Polygonum), winged or unwinged, usually 2–3-gonous, sometimes discoid, biconvex, or spheroidal, rarely 4-gonous. |
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Seeds | embryo usually straight or curved, rarely folded. |
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2n | = 40. |
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Rumex obtusifolius |
Polygonaceae subfam. polygonoideae |
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Phenology | Flowering late spring–early fall. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Waste places, roadsides, fields, shores, meadows, wet woods, swamps | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0-2300 m (0-7500 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; TN; TX; UT; VA; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; SPM; Greenland; Europe; w Asia [Introduced in North America; introduced elsewhere]
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Mainly temperate regions of North America |
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Discussion | Rumex obtusifolius, a member of subsect. Obtusifolii Rechinger f. (K. H. Rechinger 1937), is a polymorphic species represented in Eurasia by three or four rather distinct races often treated by European authors as subspecies or varieties. These taxa differ mostly in inner tepal dentation and geographic distribution. In North America the morphotypes often intergrade. In Eurasia this species is differentiated into predominantly western subsp. obtusifolius [including R. obtusifolius subsp. agrestis (Fries) Danser], eastern subsp. sylvestris (Wallroth) Rechinger f., intermediate central European subsp. transiens (Simonkai) Rechinger f., and montane subsp. subalpinus (Schur) Simonkai. Only subspp. obtusifolius and sylvestris occur in North America; the former seems to be more common. Subspecies obtusifolius differs from subsp. sylvestris in having larger and more prominently dentate inner tepals with one tubercle, or with three distinctly unequal tubercles; in subsp. sylvestris the teeth are usually less than 0.6 mm, developing only near the base of the inner tepals, and the tubercles often almost subequal. Rumex obtusifolius may be expected elsewhere in the Great Plains region of the United States and Canada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 28, species ca. 850 (16 genera, 160 species in the flora). Morphological (K. Haraldson 1978; L.-P. Ronse Decraene and J. R. Akeroyd 1988; Ronse Decraene et al. 2000; Hong S. P. et al. 1998) and molecular (A. S. Lamb Frye and K. A. Kron 2003) data provide support for separation of Persicaria from Polygonum. Further studies are needed to elucidate the relationships of allied genera, particularly Aconogonon, Bistorta, and Koenigia with Persicaria, and Fallopia and Polygonella with Polygonum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 526. | FNA vol. 5, p. 479. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Rumex | Polygonaceae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | R. crispatulus, R. rugelii | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 1: 335. (1753) | Eaton: Bot. Dict. ed. 4, 30. (1836) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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