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bristle dock, golden dock, maritime dock, rumex maritime

swamp dock

Habit Plants annual, rarely biennial [perennial], glabrous or very weakly short-papillose, mostly in inflorescence and on leaf blades, with fusiform, vertical rootstock. Plants perennial, glabrous or nearly so, with vertical rootstock.
Stems

erect (some dwarf alluvial forms may be ascending or almost prostrate), branched in distal 2/3, occasionally almost near base, (5–)15–75(–100) cm.

erect or, rarely, ascending, simple or producing axillary shoots below 1st-order inflorescence or at proximal nodes, 40–100(–150) cm.

Leaves

ocrea deciduous to partially persistent at maturity;

blade lanceolate or lanceolate-linear, rarely oblong-lanceolate, usually very gradually narrowed at both ends, (4–)7–25(–40) × (1–)1.5–4(–5) cm, at least 4 times as long as wide, base narrowly cuneate, very rarely broadly cuneate, margins entire, flat or rarely weakly undulate, apex acute, very rarely subobtuse.

blades with lateral veins forming angle of 45–60° with midvein, linear-lanceolate or lanceolate, 5–30(–40) × 1–5 cm, usually 5–7(–10) times as long as wide, normally rather thin or at most subcoriaceous, base narrowly cuneate, margins entire, flat or slightly undulate, apex acute or acuminate.

Inflorescences

terminal, occupying distal 1/2 of stem, occasionally most of stem, golden or greenish yellow, usually rather dense or interrupted in proximal part, broadly paniculate, branches spreading.

terminal and axillary, terminal usually occupying distal 1/3–1/2 of stem, usually lax, interrupted at least in basal 1/2, narrowly paniculate.

Pedicels

articulated near base or at least in proximal 1/3, filiform, 3–8 mm, articulation weakly evident.

articulated in proximal part, distinctly thickened distally, 10–17 mm, (2.5–)3–5 times as long as inner tepals, articulation distinctly or slightly swollen.

Flowers

15–30 (occasionally more) in rather dense whorls;

inner tepals narrowly triangular or narrowly rhombic-triangular, 2.5–3(–3.5) × 0.75–1.2(–1.5) mm (excluding teeth), normally ca. 2 times as long as wide, base truncate or broadly cuneate, margins prominently dentate, apex acute, very rarely subacute, straight, teeth 2–3(–4), normally at each side of margins, subulate-filiform, bristlelike, 1–3.5 mm, usually 1.5–2 times as long as width of inner tepals;

tubercles 3, equal or subequal, apex acute or subacute, smooth.

10–15(–25) in remote whorls;

inner tepals ovate-triangular or ovate-deltoid, 3.5–5 × 2.5–4 mm, longer than wide or, very rarely, as long as wide, base truncate or rounded, margins entire or, rarely, very indistinctly erose, apex acute or subacute (then with broadly triangular-lingulate tip);

tubercles 3, equal or subequal, minutely punctate and/or transversely rugose (wrinkled) in proximal part.

Achenes

light brown, small, 0.9–1.75 × 0.6–1 mm.

brown or dark brown, 2.3–3.1 × 1.6–2.2 mm.

2n

= 40.

= 60.

Rumex maritimus

Rumex verticillatus

Phenology Flowering summer–early fall. Flowering spring–early summer.
Habitat Alluvial, riparian, and coastal habitats, mostly as a ruderal species Swamps, bogs, marshes, wet meadows, irrigation ditches, wet alluvial woods
Elevation 0-500 m (0-1600 ft) 0-800 m (0-2600 ft)
Distribution
from FNA
AK; MA; NJ; YT; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

This Eurasian species is known as a casual alien from several localities in North America. Its distribution is poorly known due to confusion with native American species of this aggregate. Plants from Alaska and Yukon reported by E. Hultén (1968) as Rumex maritimus need additional study; they may be conspecific with some eastern Asian races of the R. maritimus aggregate. It is rare or almost absent in eastern Asia, where it is replaced by closely related taxa.

Species of the Rumex maritimus aggregate can be placed in a separate subsection Maritimi Rechinger f. (K. H. Rechinger 1937) or even section Orientales A. I. Baranov & B. V. Skvortzov (see A. E. Borodina 1977).

In addition to characters mentioned in the key and descriptions, additional distinctive features of Rumex maritimus are the smooth tubercles (occasionally finely striate or indistinctly pitted in herbarium specimens), and golden yellow or greenish yellow mature inflorescences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Rumex verticillatus for New Mexico (W. C. Martin and C. R. Hutchins 1980) need confirmation. The species was reported erroneously from Colorado (S. L. O’Kane et al. 1988) as a result of misidentification of R. fueginus (see W. A. Weber and R. C. Wittmann 1992).

I have not seen specimens of Rumex verticillatus from Delaware, Maine, New Hampshire, and New Jersey, but the species probably occurs in those states.

The following two species are closely related to Rumex verticillatus and sometimes treated as subspecies of it.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 529. FNA vol. 5, p. 505.
Parent taxa Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Rumex Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Axillares
Sibling taxa
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. arcticus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hastatulus, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. venosus, R. verticillatus, R. violascens
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. arcticus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hastatulus, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. maritimus, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. venosus, R. violascens
Synonyms Lapathum minus, R. aureus
Name authority Linnaeus: Sp. Pl. 1: 335. (1753) Linnaeus: Sp. Pl. 1: 334. (1753)
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