Rumex hymenosepalus |
Rumex arcticus |
|
---|---|---|
Arizona dock, canaigre, canaigre dock, cañaigre dock, wild-rhubarb |
arctic dock |
|
Habit | Plants perennial, glabrous or indistinctly papillose-pubescent, with distinctly tuberous roots and short rhizomes. | Plants perennial, glabrous or nearly so, with fusiform, oblique rootstock, occasionally with horizontal, short-creeping rhizome. |
Stems | usually erect, rarely ascending, branched above middle, 25–90(–100) cm. |
erect, simple or branched in distal 2/3 (then with few, comparatively short branches), 10–70(–100) cm. |
Leaves | ocrea prominent and persistent at maturity, whitish or silvery white, membranous; blade oblong, oblong-elliptic, or obovate-lanceolate, (5–)8–30 × 2–8(–12) cm, base cuneate or narrowly cuneate, margins entire, flat or indistinctly crisped, apex acute or acuminate, rarely obtuse. |
ocrea deciduous or partially persistent at maturity; blade narrowly lanceolate, lanceolate, or oblong-lanceolate, normally 5–15(–20) × 1.5–5 cm, base cuneate to broadly cuneate, rarely truncate or very weakly cordate, margins entire or rarely indistinctly repand, flat, apex acute or subacute. |
Inflorescences | terminal, occupying distal 1/2 of stem, narrowly paniculate, rarely simple. |
terminal, occupying distal 1/2–2/3 of stem, interrupted, paniculate, simple or nearly so (branches, when present usually less than 7–8 cm). |
Pedicels | articulated near middle or in proximal 1/3, filiform, 5–15(–20) mm, articulation indistinct. |
articulated in proximal 1/3, filiform, 5–13(–17) mm, usually not more than 2–2.5 times as long as inner tepals, articulation weakly evident, not swollen. |
Flowers | 5–20 in whorls; inner tepals oblong-cordate or orbiculate-cordate, 11–16 × 9.5–14 mm, base sinuate or emarginate, margins entire, rarely with few extremely small denticles at base, apex obtuse or subacute; tubercles absent. |
7–15 in whorls; inner tepals ovate, 4.5–7.5(–8) × 4–6(–7) mm, base truncate to weakly cordate, margins entire, apex obtuse or subacute; tubercles absent. |
Achenes | brown or reddish brown, 4–5(–7) × 2.5–4.5(–5) mm. |
reddish brown, 3–4 × 1.5–2 mm. |
2n | = 40. |
= 40, 120, ca. 170, ca. 200. |
Rumex hymenosepalus |
Rumex arcticus |
|
Phenology | Flowering spring. | Flowering late spring–summer. |
Habitat | Sandy and rocky places: plains, slopes, stream beds, alkaline soils | Moist tundra, marshes, river valleys, sandy and gravelly shores and slopes |
Elevation | 0-1700(-2000) m (0-5600(-6600) ft) | 0-1000 m (0-3300 ft) |
Distribution |
AZ; CA; CO; MT; NM; NV; OK; TX; UT; WY; Mexico (Baja California, Chihuahua)
|
AK; BC; NT; NU; YT; ne Europe; n Asia (arctic and subarctic zones) |
Discussion | Rumex hymenosepalus is the only species of subsect. Hymenosepali Rechinger f. Two varieties have been recognized. The typical variety has achenes 5 mm and ovate-elliptic or oblong-cordate inner tepals with a subacute apex. Variety salinus (A. Nelson) Rechinger f. has larger achenes (to 7 mm) and almost orbiculate inner tepals with an obtuse apex. Rumex hymenosepalus was reported also from Montana (J. E. Dawson 1979), but no exact localities were given. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Rumex arcticus is polymorphic, as are R. aquaticus in the narrow sense and R. occidentalis. However, unlike R. aquaticus and R. occidentalus, it is represented by at least two chromosome races (G. A. Mulligan and C. Frankton 1972; Á. Löve 1986). Plants morphologically transitional between R. arcticus and R. aquaticus were described from Kamchatka as R. kamtschadalus (= R. arcticus var. kamtschadalus). The same forms occasionally occur in northwestern North America. According to Á. Löve and D. Löve (1975b) and Á. Löve (1986), they are usually tetraploids (2n = 40) and deserve recognition at the species level. However, they are not always morphologically distinct from R. arcticus. The group needs additional study, and at present I prefer to keep the tetraploid plants provisionally within R. arcticus, regarding them as var. kamtschadalus. A few highly sterile specimens with mostly abortive flowers, which I have seen in Alaskan herbarium material, most probably represent hybrids between tetraploid and 12-ploid races of the R. aquaticus group. Plants with unusually wide, triangular-oblong, or almost ovate leaves were described as var. latifolius Tolmatchew. This seems to be a predominant variety on the Beringian coast of Chukotka and Wrangel and Ratmanov islands (A. I. Tolmachew 1966). I also have seen at least two collections of this variety from the western coast of Alaska. The enigmatic var. perlatus Hultén may belong here (see discussion under 8. R. lapponicus). Rumex arcticus has been reported from Churchill in northeastern Manitoba (H. J. Scoggan 1978–1979, part 3). That record needs confirmation because some northern forms of R. arcticus and R. occidentalis are similar. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 515. | FNA vol. 5, p. 518. |
Parent taxa | Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Rumex | Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Rumex |
Sibling taxa | ||
Synonyms | R. arizonicus, R. hymenosepalus var. salinus, R. salinus, R. saxei | R. aquaticus subsp. arcticus, R. arcticus var. kamtschadalus, R. arcticus var. latifolius, R. domesticus var. nanus, R. kamtschadalus, R. longifolius var. nanus, R. ursinus |
Name authority | Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 177. (1859) | Trautvetter: in A. T. von Middendorff, Reise Siber. 1(2,1): 29. (1847) |
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