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heartwing dock, heartwing sorrel, wild dock, wild sorrel

swamp dock

Habit Plants annual or short-lived perennial, glabrous, with vertical rootstock. Plants perennial, glabrous or nearly so, with vertical rootstock.
Stems

solitary or several from base, erect or ascending, branched in distal 2/3 (in inflorescence), 10–40(–45) cm.

erect or, rarely, ascending, simple or producing axillary shoots below 1st-order inflorescence or at proximal nodes, 40–100(–150) cm.

Leaves

blade obovate-oblong, ovate-lanceolate, oblong-lanceolate, or lanceolate; 2–6(–10) × 0.5–2 cm, base hastate (with spreading lobes), auriculate, or occasionally without evident lobes, margins entire, flat, apex obtuse or subacute.

blades with lateral veins forming angle of 45–60° with midvein, linear-lanceolate or lanceolate, 5–30(–40) × 1–5 cm, usually 5–7(–10) times as long as wide, normally rather thin or at most subcoriaceous, base narrowly cuneate, margins entire, flat or slightly undulate, apex acute or acuminate.

Inflorescences

terminal, occupying distal 2/3 of stem, usually lax and interrupted, narrowly paniculate.

terminal and axillary, terminal usually occupying distal 1/3–1/2 of stem, usually lax, interrupted at least in basal 1/2, narrowly paniculate.

Pedicels

articulated in proximal part, filiform, 1.5–2.5(–3) mm, articulation indistinct or slightly swollen.

articulated in proximal part, distinctly thickened distally, 10–17 mm, (2.5–)3–5 times as long as inner tepals, articulation distinctly or slightly swollen.

Flowers

3–6(–8) in whorls;

inner tepals orbiculate or broadly ovate, 2.5–3.2 × 2.7–3.2 mm, base broadly cordate or rounded, apex obtuse or subacute;

tubercles absent or some inner tepals with slightly swollen central veins.

10–15(–25) in remote whorls;

inner tepals ovate-triangular or ovate-deltoid, 3.5–5 × 2.5–4 mm, longer than wide or, very rarely, as long as wide, base truncate or rounded, margins entire or, rarely, very indistinctly erose, apex acute or subacute (then with broadly triangular-lingulate tip);

tubercles 3, equal or subequal, minutely punctate and/or transversely rugose (wrinkled) in proximal part.

Achenes

brown or dark brown, 0.9–1.2 × 0.6–0.8 mm.

brown or dark brown, 2.3–3.1 × 1.6–2.2 mm.

2n

= 8 (pistillate plants), 9 (staminate plants), 10 (both sexes).

= 60.

Rumex hastatulus

Rumex verticillatus

Phenology Flowering spring–summer. Flowering spring–early summer.
Habitat Dry to moist alluvial and ruderal habitats, river valleys, sandy plains, meadows, waste places Swamps, bogs, marshes, wet meadows, irrigation ditches, wet alluvial woods
Elevation 0-500 m (0-1600 ft) 0-800 m (0-2600 ft)
Distribution
from FNA
AL; AR; FL; GA; IL; IN; KS; LA; MD; MO; MS; NC; NY; OK; PA; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Rumex hastatulus is distinct in subg. Acetosa and belongs to the monotypic subsect. Americanae Á. Löve & N. Sarkar. It is represented by at least two chromosome races: populations occurring from North Carolina to Florida and Mississippi normally have 2n = 8 in pistillate plants and 2n = 9 in staminate plants; populations from Louisiana to Texas and Oklahoma predominantly have 2n = 10 in both sexes. Rumex hastatulus has been reported from New Mexico (W. C. Martin and C. R. Hutchins 1980, vol. 1), but those records need confirmation. When fruiting, R. hastatulus has large inner tepals that distinguish it from R. acetosella, with which it is occasionally confused.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Rumex verticillatus for New Mexico (W. C. Martin and C. R. Hutchins 1980) need confirmation. The species was reported erroneously from Colorado (S. L. O’Kane et al. 1988) as a result of misidentification of R. fueginus (see W. A. Weber and R. C. Wittmann 1992).

I have not seen specimens of Rumex verticillatus from Delaware, Maine, New Hampshire, and New Jersey, but the species probably occurs in those states.

The following two species are closely related to Rumex verticillatus and sometimes treated as subspecies of it.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 502. FNA vol. 5, p. 505.
Parent taxa Polygonaceae > subfam. Polygonoideae > Rumex > subg. Acetosa Polygonaceae > subfam. Polygonoideae > Rumex > subg. Rumex > sect. Axillares
Sibling taxa
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. arcticus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. maritimus, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. venosus, R. verticillatus, R. violascens
R. acetosa, R. acetosella, R. alpinus, R. altissimus, R. arcticus, R. beringensis, R. britannica, R. brownii, R. bucephalophorus, R. californicus, R. chrysocarpus, R. confertus, R. conglomeratus, R. crassus, R. crispus, R. cristatus, R. cuneifolius, R. densiflorus, R. dentatus, R. ellipticus, R. fascicularis, R. floridanus, R. fueginus, R. graminifolius, R. hastatulus, R. hesperius, R. hymenosepalus, R. kerneri, R. krausei, R. lacustris, R. lapponicus, R. longifolius, R. maritimus, R. mexicanus, R. nematopodus, R. obovatus, R. obtusifolius, R. occidentalis, R. orthoneurus, R. pallidus, R. palustris, R. paraguayensis, R. patientia, R. paucifolius, R. persicarioides, R. praecox, R. pseudonatronatus, R. pulcher, R. pycnanthus, R. salicifolius, R. sanguineus, R. sibiricus, R. spiralis, R. stenophyllus, R. subarcticus, R. thyrsiflorus, R. tomentellus, R. transitorius, R. triangulivalvis, R. utahensis, R. venosus, R. violascens
Synonyms Acetosa hastatula, R. engelmannii
Name authority Baldwin: in S. Elliott, Sketch Bot. S. Carolina 1: 416. (1817) Linnaeus: Sp. Pl. 1: 334. (1753)
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