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cone-flower, rudbeckia
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brown-eyed susan
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| Annuals, perennials, subshrubs, or shrubs [perennials], mostly 10–80(–200) cm. |
Annuals, biennials, or perennials, to 100(–200) cm (rhizomatous, fibrous rooted, or taprooted). |
erect, branched distally or ± throughout. |
green. |
cauline; mostly opposite (distal sometimes alternate); petiolate or sessile; blades mostly lanceolate to oblanceolate overall, usually 1–3-pinnately lobed or -pinnatisect, ultimate margins toothed or entire, faces glabrous or hairy (oil-glands scattered and/or submarginal). |
green, not glaucous; basal (usually withering before flowering) usually petiolate, blades linear, lanceolate to ovate, or elliptic to oblanceolate, sometimes lobed, bases attenuate, cuneate, or rounded, margins entire, dentate, or serrate, apices acute to obtuse, faces glabrous or hairy, sometimes gland-dotted; cauline usually petiolate (at least proximal), distal usually sessile, blades linear or ovate to pandurate, sometimes lobed, lobes 3(–5), bases auriculate, attenuate, or cordate, margins entire or serrate, apices acute to obtuse, faces glabrous or hairy, sometimes gland-dotted. |
narrowly cylindric or fusiform to turbinate or broadly campanulate, 1–12+ mm diam. |
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convex to conic, smooth or finely pitted, epaleate. |
usually conic to hemispheric, rarely columnar; paleae surpassing cypselae, margins sometimes ciliate, apices obtuse or rounded to acute or attenuate to apiculate, faces glabrous or hairy abaxially. |
0 or 1–8(–13+) (to 100+ in “double” cultivars), pistillate, fertile (except “double” cultivars); corollas yellow or orange, red-brown (with or without yellow/orange), or white. |
6–25+; corollas usually yellow-orange proximally, yellow distally, sometimes with basal maroon splotch (orangish red to maroon in R. graminifolia). |
6–120+, bisexual, fertile; corollas greenish yellow to orange, sometimes tipped with red or red-brown, tubes much longer than or about equaling funnelform throats, lobes 5, deltate to lance-linear (equal or 2 sinuses deeper than others). |
50–800+; corollas proximally yellow to yellowish green, distally usually brown-purple, lobes sometimes yellowish or greenish; anther appendages sometimes gland-dotted; styles 3–5+ mm, branches 1–1.8 mm, proximal 1/2–4/5 stigmatic, apices acute to rounded. |
persistent, 3–21+ in 1–2 series (connate to 7/8+ their lengths, usually streaked and/or dotted with oil-glands). |
in 2(–3) series. |
0. |
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radiate or discoid, borne singly or in ± corymbiform arrays. |
borne singly or in loose, corymbiform to paniculiform arrays. |
narrowly obpyramidal or fusiform-terete, sometimes weakly flattened, glabrous or hairy; pappi persistent, of 2–5(–10) dissimilar, distinct or connate scales in ± 1 series: 0–5+ oblong to lanceolate, erose-truncate or laciniate plus 0–2(–5) longer, subulate to aristate. |
1.5–3.5(–4) mm; pappi usually coroniform (sometimes cypselae each with glandular hairs around apices in R. heliopsidis). |
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8–30 × 5–25 mm. |
= 12. |
= 19. |
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North America [Introduced in Europe] |
e North America |
Species 23 (23 in the flora). The species of Rudbeckia are distributed among three major clades or lineages. Although relationships among the lineages are not robustly resolved, the lineages are treated here as sections (as they have been traditionally). Rudbeckia hirta and sometimes other species of the genus are used in experimental studies relating to initiation of flowering and hairy root culture. Most species are rich sources of phytochemicals that may offer potential for pharmaceutical or other uses. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 9 (9 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae |
Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia |
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1. Annuals (stems glabrous); leaves sessile, blades elliptic, lanceolate, oblanceolate, oblong, or ovate (not lobed, bases auriculate and clasping, faces glabrous, glaucous); phyllaries in 2 series (lengths of outer 2–4+ times inner, inner sometimes interpreted as paleae); paleae surpassing cypselae, margins ciliate, faces usually glabrous; pappi 0 | sect. Dracopis |
1. Annuals, biennials, or perennials (if epappose annuals, stems hairy); leaves petiolate or sessile, blades elliptic, lanceolate, linear, oblanceolate, ovate, or spatulate (often lobed, faces glabrous or hairy, sometimes glaucous; if leaves sessile and not lobed and bases auriculate and clasping, then robust perennials); phyllaries in 2(–3) series; paleae sometimes surpassing cypselae, margins usually ciliate, rarely eciliate, faces glabrous or hairy; pappi usually coroniform or of 2–6+ scales, sometimes 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis) | → 2 |
2. Leaves often bluish green, glaucous; receptacles usually conic to columnar (paleae not surpassing cypselae, except in R. laciniata: paleae surpassing cypselae, receptacles sometimes hemispheric to ovoid, disc corolla lobes yellow); ray florets 0 or 8–15+, corollas bright yellow; disc corollas proximally yellow to yellowish green, distally yellow or greenish to brown-purple; cypselae (3–)3.5–7.5 mm; pappi coroniform, or of 2–6 scales 0.1–2.5 mm | sect. Macrocline |
2. Leaves green, not glaucous; receptacles usually conic to hemispheric, rarely columnar (paleae surpassing cypselae); ray florets 6–25+, corollas usually yellow-orange proximally, yellow distally, sometimes with basal maroon splotch (orangish red to maroon in R. graminifolia); disc corollas proximally yellow to yellowish green, distally usually brown-purple, lobes sometimes yellowish or greenish; cypselae 1.5–3.5(–4) mm; pappi coroniform, or of 8+ unequal scales 0.1–2 mm, or 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis) | sect. Rudbeckia |
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1. Leaves 0.2–1 cm wide (grasslike, lengths 10+ times widths); rays orangish red to maroon; Florida panhandle | R. graminifolia |
1. Leaves (0.4–)1–15 cm wide (if less than 1 cm wide, lengths less than 10 times widths); rays yellow to yellow-orange (at least distally); e North America | → 2 |
2. Annuals, biennials, or perennials (taprooted or roots fibrous); pappi 0 or coroniform (to 0.1 mm) | → 3 |
2. Perennials (usually rhizomatous and/or stoloniferous, roots fibrous, caudices sometimes woody); pappi 0 or coroniform (0.1–0.5 mm; sometimes cypselae each with glandular hairs around apices in R. heliopsidis) | → 4 |
3. Stems and leaves coarsely hispid to hirsute; style branch apices subulate; e NorthAmerica | R. hirta |
3. Stems and leaves softly pilose to woolly; style branch apices acute to obtuse; Alabama, Florida, Georgia, South Carolina | R. mollis |
4. Proximal cauline leaves elliptic or ovate, usually 3(–5)-lobed; paleae cuspidate (tips awnlike, 1.5+ mm) | R. triloba |
4. Proximal cauline leaves elliptic, linear, spatulate, or ovate, rarely lobed (3–5–lobed in R. subtomentosa); paleae acute, obtuse, or rounded | → 5 |
5. Stems and leaves densely hirsute; proximal cauline leaves 3(–5)–lobed; heads (usually 8–25) in loose, corymbiform to paniculiform arrays | R. subtomentosa |
5. Stems and leaves glabrous or sparsely to moderately hairy; proximal cauline leaves not lobed; heads borne singly or (2–12) in corymbiform arrays | → 6 |
6. Leaves: abaxial faces glabrous or hirsute to strigose (not gland-dotted); paleae: abaxial tips usually glabrous, rarely pilose | → 7 |
6. Leaves: abaxial faces glabrous or strigose and gland-dotted; paleae: abaxial tips canescent or strigose, sometimes gland-dotted as well | → 8 |
7. Plants stoloniferous (rosettes forming at stolon apices; stem branches spreading); basal leaves lanceolate to broadly ovate or elliptic | R. fulgida |
7. Plants not stoloniferous (rosettes at bases of aerial stems; stem branches ascending); basal leaves linear to narrowly spatulate | R. missouriensis |
8. Leaf blades elliptic, lanceolate, or ovate (flat); heads usually (4–8) in ± corymbiform arrays, sometimes borne singly; rays 6–12 (± spreading); pappi coroniform, to 1.5 mm (sometimes cypselae each with glandular hairs around apices); Alabama, Georgia, North Carolina, South Carolina, Virginia | R. heliopsidis |
8. Leaf blades elliptic, lanceolate, or ovate (± conduplicate); heads usually borne singly; rays 12–25 (reflexed); pappi coroniform, to 0.5 mm; midwestern and sc United States | R. grandiflora |
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| Linnaeus: Sp. Pl. 2: 906. (1753): Gen. Pl. ed. 5, 387. (1754) |
unknown |
| FNA vol. 21, p. 44. Treatment authors: Lowell E. Urbatsch, Patricia B. Cox. |
FNA vol. 21, p. 52. Treatment authors: Lowell E. Urbatsch, Patricia B. Cox. |
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