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cone-flower, rudbeckia

sweet coneflower

Habit Annuals, perennials, subshrubs, or shrubs [perennials], mostly 10–80(–200) cm. Annuals, 25–60(–120) cm (taprooted).
Stems

erect, branched distally or ± throughout.

green (glaucous, glabrous).

Leaves

cauline; mostly opposite (distal sometimes alternate);

petiolate or sessile;

blades mostly lanceolate to oblanceolate overall, usually 1–3-pinnately lobed or -pinnatisect, ultimate margins toothed or entire, faces glabrous or hairy (oil-glands scattered and/or submarginal).

bluish green, glaucous;

basal (seldom persisting to flowering) petiolate or sessile, blades elliptic, lanceolate, oblanceolate, oblong, or ovate, not lobed, bases (cauline) auriculate and clasping, margins crenate, entire or serrate, apices acute or acuminate, faces glabrous;

cauline sessile, blades elliptic, lanceolate, oblanceolate, oblong, or ovate, bases auriculate and clasping, margins crenate, entire or serrate, apices acute or acuminate, faces glabrous.

Involucres

narrowly cylindric or fusiform to turbinate or broadly campanulate, 1–12+ mm diam.

Receptacles

convex to conic, smooth or finely pitted, epaleate.

ovoid to conic;

paleae surpassing cypselae (margins ciliate), apices obtuse to acute, often mucronate, faces hairy subapically, glabrous near apices.

Ray florets

0 or 1–8(–13+) (to 100+ in “double” cultivars), pistillate, fertile (except “double” cultivars);

corollas yellow or orange, red-brown (with or without yellow/orange), or white.

6–10+;

corollas yellow, sometimes partly orange or maroon (laminae often with proximal maroon splotch).

Disc florets

6–120+, bisexual, fertile;

corollas greenish yellow to orange, sometimes tipped with red or red-brown, tubes much longer than or about equaling funnelform throats, lobes 5, deltate to lance-linear (equal or 2 sinuses deeper than others).

to 400+;

corollas proximally greenish yellow, distally purplish;

styles ca. 5 mm, branches ca. 1.7 mm, proximal 1/2 stigmatic, apices subulate.

Phyllaries

persistent, 3–21+ in 1–2 series (connate to 7/8+ their lengths, usually streaked and/or dotted with oil-glands).

in 2 series (lengths of outer 2–5+ times inner, inner similar to and sometimes interpreted as paleae).

Calyculi

0.

Heads

radiate or discoid, borne singly or in ± corymbiform arrays.

in loose, corymbiform arrays or borne singly.

Cypselae

narrowly obpyramidal or fusiform-terete, sometimes weakly flattened, glabrous or hairy;

pappi persistent, of 2–5(–10) dissimilar, distinct or connate scales in ± 1 series: 0–5+ oblong to lanceolate, erose-truncate or laciniate plus 0–2(–5) longer, subulate to aristate.

1.8–2.5 mm;

pappi 0.

Discs

15–30 × 8–15 mm.

x

= 12.

= 16.

Rudbeckia

Rudbeckia sect. Dracopis

Distribution
from USDA
North America [Introduced in Europe]
[BONAP county map]
c United States; e United States
Discussion

Species 23 (23 in the flora).

The species of Rudbeckia are distributed among three major clades or lineages. Although relationships among the lineages are not robustly resolved, the lineages are treated here as sections (as they have been traditionally). Rudbeckia hirta and sometimes other species of the genus are used in experimental studies relating to initiation of flowering and hairy root culture. Most species are rich sources of phytochemicals that may offer potential for pharmaceutical or other uses.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 1.

Rudbeckia sect. Dracopis is sometimes recognized at generic rank as Dracopis. DNA-based phylogenetic analyses support its placement in Rudbeckia, where it was treated by Gray as a section. The single species resembles those in sect. Macrocline; it is often placed there based on DNA evidence; the relationship is not uniformly conclusive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia
Subordinate taxa
R. sect. Dracopis, R. sect. Macrocline, R. sect. Rudbeckia
Key
1. Annuals (stems glabrous); leaves sessile, blades elliptic, lanceolate, oblanceolate, oblong, or ovate (not lobed, bases auriculate and clasping, faces glabrous, glaucous); phyllaries in 2 series (lengths of outer 2–4+ times inner, inner sometimes interpreted as paleae); paleae surpassing cypselae, margins ciliate, faces usually glabrous; pappi 0
sect. Dracopis
1. Annuals, biennials, or perennials (if epappose annuals, stems hairy); leaves petiolate or sessile, blades elliptic, lanceolate, linear, oblanceolate, ovate, or spatulate (often lobed, faces glabrous or hairy, sometimes glaucous; if leaves sessile and not lobed and bases auriculate and clasping, then robust perennials); phyllaries in 2(–3) series; paleae sometimes surpassing cypselae, margins usually ciliate, rarely eciliate, faces glabrous or hairy; pappi usually coroniform or of 2–6+ scales, sometimes 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis)
→ 2
2. Leaves often bluish green, glaucous; receptacles usually conic to columnar (paleae not surpassing cypselae, except in R. laciniata: paleae surpassing cypselae, receptacles sometimes hemispheric to ovoid, disc corolla lobes yellow); ray florets 0 or 8–15+, corollas bright yellow; disc corollas proximally yellow to yellowish green, distally yellow or greenish to brown-purple; cypselae (3–)3.5–7.5 mm; pappi coroniform, or of 2–6 scales 0.1–2.5 mm
sect. Macrocline
2. Leaves green, not glaucous; receptacles usually conic to hemispheric, rarely columnar (paleae surpassing cypselae); ray florets 6–25+, corollas usually yellow-orange proximally, yellow distally, sometimes with basal maroon splotch (orangish red to maroon in R. graminifolia); disc corollas proximally yellow to yellowish green, distally usually brown-purple, lobes sometimes yellowish or greenish; cypselae 1.5–3.5(–4) mm; pappi coroniform, or of 8+ unequal scales 0.1–2 mm, or 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis)
sect. Rudbeckia
Synonyms Obeliscaria subg. Dracopis, section Dracopis
Name authority Linnaeus: Sp. Pl. 2: 906. (1753): Gen. Pl. ed. 5, 387. (1754) (Cassini) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 1(2): 263. (1884)
Source FNA vol. 21, p. 44. Treatment authors: Lowell E. Urbatsch, Patricia B. Cox. FNA vol. 21, p. 45. Treatment authors: Lowell E. Urbatsch, Patricia B. Cox.
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