California coneflower, waxy cone-flower
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Perennials, to 150 cm (roots fibrous). |
Perennials, 50–300 cm (robust, rhizomatous or fibrous rooted). |
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green, bluish green, or purplish (usually glaucous). |
bluish green (heavily glaucous), blades lanceolate to elliptic (not lobed), leathery, bases attenuate, margins entire or remotely serrulate, apices acute, faces glabrous; basal petiolate, 20–50 × 4–10 cm; cauline petiolate or sessile, 10–25 × 2–8 cm. |
often bluish green, glaucous; basal (usually persistent to flowering) petiolate or sessile; blades linear, lanceolate to ovate, or oblong to elliptic, usually lobed, bases attenuate, cuneate, or rounded, ultimate margins entire, crenate, dentate, or serrate, apices acute to obtuse, faces glabrous or hairy, sometimes glaucous; cauline petiolate or sessile, blades linear or ovate to pandurate, sometimes lobed, bases auriculate, rounded, or attenuate, ultimate margins entire or dentate to serrate, apices acute to obtuse, faces glabrous or hairy, sometimes glaucous. |
conic to columnar; paleae 4–6.5 mm, apices acute, often attenuate, abaxial tips hairy. |
usually conic to columnar (hemispheric to ovoid in R. laciniata); paleae not surpassing cypselae (except in R. laciniata), margins ciliate, apices acute to obtuse or rounded to truncate, attenuate to apiculate, usually hairy. |
7–15; laminae elliptic to oblong, 25–40 × 8–14 mm, abaxially hairy. |
0 or 8–15+; corollas bright yellow. |
250–400+; corollas yellowish green, 3–4 mm; style branches ca. 1 mm, apices acute. |
(100–)150–300(–600+); corollas proximally yellow to yellowish green, distally yellow or greenish to brown-purple; anther appendages sometimes abaxially gland-dotted; styles 3–7 mm, branches 1–2.2 mm, proximal 2/3–4/5 stigmatic, apices acute to rounded. |
to 1.5 cm. |
in 2(–3) series. |
borne singly or (2–10) in ± corymbiform arrays. |
borne singly or in loose, corymbiform to paniculiform arrays. |
4–5.5 mm; pappi coroniform or of ± connate scales, to 1.2 mm. |
(3–)3.5–7.5 mm; pappi usually coroniform or of 2–6, unequal scales, 0.1–2.5 mm, sometimes 0. |
15–35 × 14–22 mm. |
12–60(–80) × 10–30 mm. |
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= 18. |
= 36. |
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Flowering summer–fall. |
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Meadows, seeps, streamsides |
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60–1300 m (200–4300 ft) |
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CA; OR
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North America |
Rudbeckia glaucescens often grows on serpentine and often with Darlingtonia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 13 (13 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Basal and cauline leaves 0.2–1.5 cm wide | R. mohrii |
1. Basal and cauline leaves 2–15(–45) cm wide | → 2 |
2. Blades of basal and proximal cauline leaves elliptic, lanceolate, or ovate, usually 1–2-pinnatifid or -pinnately compound (leaflets/lobes 3–11, distal cauline leaves sometimes lobed as well) | → 3 |
2. Blades of basal, and proximal and mid, cauline leaves elliptic or lanceolate (not lobed, sometimes coarsely toothed, margins of distal cauline leaves crenate, dentate, entire, or toothed, not lobed) | → 7 |
3. Blades of all but distalmost leaves usually 1–2-pinnatifid or pinnately lobed (basal leaves not lobed and proximal cauline usually 3-lobed in var. heterophylla, known only from Florida); receptacles hemispheric or globose to ovoid; disc corolla lobes yellow (rhizomes elongate, slender, plants usually colonial); Rocky Mountains and e United States | R. laciniata |
3. Blades of basal leaves ± pinnatifid to pinnately lobed (distal leaves sometimes not lobed); receptacles columnar, conic, cylindric, or ovoid; disc corolla lobes yellowish green, greenish, or purplish (rhizomes stout, plants usually not colonial); w United States (including Rocky Mountains) | → 4 |
4. Leaves sparsely to densely hairy on abaxial or both faces | → 5 |
4. Leaves usually glabrous, sometimes sparsely hairy on veins abaxially | → 6 |
5. Leaves moderately to densely hairy (both faces); rays 0; pappi of 4 scales, to 1 mm; Washington | R. alpicola |
5. Leaves sparsely hairy (abaxial faces); rays 8–21; pappi coroniform or of scales, 0.5–1.5 mm; California | R. californica |
6. Rays 7–15; phyllaries to 1.5 cm (apices acute to rounded); discs 15–35 mm; California | R. klamathensis |
6. Rays 0; phyllaries to 4 cm (apices attenuate); discs 20–60 mm; Colorado Rockies, Utah | R. montana |
7. Mid (and often proximal) cauline leaves sessile, bases of blades usually auriculate (clasping) to truncate, sometimes rounded, faces (one or both) glabrate or hairy, or glabrous | → 8 |
7. Mid cauline leaves petiolate, bases of blades attenuate to cuneate (not clasping), faces (one or both) glabrous or hairy | → 10 |
8. Leaves bluish green (fresh, heavily glaucous; dried blades white under reflected UV light, at least abaxially); discs 40–80 mm; adjoining areas of Arkansas, Louisiana,Oklahoma, Texas (introduced South Carolina) | R. maxima |
8. Leaves green (fresh; dried blades dark under reflected UV light, at least abaxially); discs 12–25 mm; Alabama, Florida, Louisiana, Texas | → 9 |
9. Cauline leaf bases auriculate, clasping; heads (usually 10+) in paniculiform ar-rays; discs 12–16 mm; paleae 4–6 mm; Alabama, Florida | R. auriculata |
9. Cauline leaf bases auriculate or rounded, not clasping; heads (usually to 15) in ± corymbiform arrays; discs 12–25 mm; paleae 6–8 mm (stems often purplish); w Louisiana, e Texas 13. Rudbeckia scabrifolia | R. scabrifolia |
10. Leaf margins serrate or entire, faces usually sparsely to densely hairy (mostly abaxially), rarely glabrous; rays 0; n Rocky Mountains and Pacific states | R. occidentalis |
10. Leaf margins entire, serrate, serrulate, or toothed, faces glabrous or sparsely hairy; rays 7–16; Pacific states or se United States | → 11 |
11. Leaves bluish green (heavily glaucous); California, s Oregon | R. glaucescens |
11. Leaves green (sometimes lightly glaucous); se United States | → 12 |
12. Basal leaf blades 15–50 × 3–9 cm (lengths ± 5 times widths); discs 20–45 mm; paleae 6–8 mm, apices acute (appressed in young heads); cypselae 5–7.5 mm; w Louisiana, e Texas | R. texana |
12. Basal leaf blades 15–60 × 2–8 cm (lengths ± 7 times widths); discs 10–30 mm; paleae 5–6 mm, apices acute to acuminate (erect to spreading in young heads); cypselae 3–5.5 mm; n Florida, s Georgia | R. nitida |
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FNA vol. 21, p. 48. |
FNA vol. 21, p. 46. |
Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia > sect. Macrocline |
Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia |
R. alpicola, R. amplexicaulis, R. auriculata, R. californica, R. fulgida, R. graminifolia, R. grandiflora, R. heliopsidis, R. hirta, R. klamathensis, R. laciniata, R. maxima, R. missouriensis, R. mohrii, R. mollis, R. montana, R. nitida, R. occidentalis, R. scabrifolia, R. subtomentosa, R. texana, R. triloba |
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R. alpicola, R. auriculata, R. californica, R. glaucescens, R. klamathensis, R. laciniata, R. maxima, R. mohrii, R. montana, R. nitida, R. occidentalis, R. scabrifolia, R. texana |
R. californica var. glauca |
R. subg. Macrocline |
Eastwood: Leafl. W. Bot. 2: 55. (1937) |
Torrey & A. Gray: Fl. N. Amer. 2: 312. (1842) |
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