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California coneflower, waxy cone-flower

cone-flower, rudbeckia

Habit Perennials, to 150 cm (roots fibrous). Annuals, perennials, subshrubs, or shrubs [perennials], mostly 10–80(–200) cm.
Stems

erect, branched distally or ± throughout.

Leaves

bluish green (heavily glaucous), blades lanceolate to elliptic (not lobed), leathery, bases attenuate, margins entire or remotely serrulate, apices acute, faces glabrous;

basal petiolate, 20–50 × 4–10 cm;

cauline petiolate or sessile, 10–25 × 2–8 cm.

cauline; mostly opposite (distal sometimes alternate); petiolate or sessile;

blades mostly lanceolate to oblanceolate overall, usually 1–3-pinnately lobed or -pinnatisect, ultimate margins toothed or entire, faces glabrous or hairy (oil-glands scattered and/or submarginal).

Involucres

narrowly cylindric or fusiform to turbinate or broadly campanulate, 1–12+ mm diam.

Receptacles

conic to columnar;

paleae 4–6.5 mm, apices acute, often attenuate, abaxial tips hairy.

convex to conic, smooth or finely pitted, epaleate.

Ray florets

7–15;

laminae elliptic to oblong, 25–40 × 8–14 mm, abaxially hairy.

0 or 1–8(–13+) (to 100+ in “double” cultivars), pistillate, fertile (except “double” cultivars);

corollas yellow or orange, red-brown (with or without yellow/orange), or white.

Disc florets

250–400+;

corollas yellowish green, 3–4 mm;

style branches ca. 1 mm, apices acute.

6–120+, bisexual, fertile;

corollas greenish yellow to orange, sometimes tipped with red or red-brown, tubes much longer than or about equaling funnelform throats, lobes 5, deltate to lance-linear (equal or 2 sinuses deeper than others).

Phyllaries

to 1.5 cm.

persistent, 3–21+ in 1–2 series (connate to 7/8+ their lengths, usually streaked and/or dotted with oil-glands).

Calyculi

0.

Heads

borne singly or (2–10) in ± corymbiform arrays.

radiate or discoid, borne singly or in ± corymbiform arrays.

Cypselae

4–5.5 mm;

pappi coroniform or of ± connate scales, to 1.2 mm.

narrowly obpyramidal or fusiform-terete, sometimes weakly flattened, glabrous or hairy;

pappi persistent, of 2–5(–10) dissimilar, distinct or connate scales in ± 1 series: 0–5+ oblong to lanceolate, erose-truncate or laciniate plus 0–2(–5) longer, subulate to aristate.

Discs

15–35 × 14–22 mm.

x

= 12.

2n

= 36.

Rudbeckia glaucescens

Rudbeckia

Phenology Flowering summer–fall.
Habitat Meadows, seeps, streamsides
Elevation 60–1300 m (200–4300 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from USDA
North America [Introduced in Europe]
[BONAP county map]
Discussion

Rudbeckia glaucescens often grows on serpentine and often with Darlingtonia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 23 (23 in the flora).

The species of Rudbeckia are distributed among three major clades or lineages. Although relationships among the lineages are not robustly resolved, the lineages are treated here as sections (as they have been traditionally). Rudbeckia hirta and sometimes other species of the genus are used in experimental studies relating to initiation of flowering and hairy root culture. Most species are rich sources of phytochemicals that may offer potential for pharmaceutical or other uses.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Annuals (stems glabrous); leaves sessile, blades elliptic, lanceolate, oblanceolate, oblong, or ovate (not lobed, bases auriculate and clasping, faces glabrous, glaucous); phyllaries in 2 series (lengths of outer 2–4+ times inner, inner sometimes interpreted as paleae); paleae surpassing cypselae, margins ciliate, faces usually glabrous; pappi 0
sect. Dracopis
1. Annuals, biennials, or perennials (if epappose annuals, stems hairy); leaves petiolate or sessile, blades elliptic, lanceolate, linear, oblanceolate, ovate, or spatulate (often lobed, faces glabrous or hairy, sometimes glaucous; if leaves sessile and not lobed and bases auriculate and clasping, then robust perennials); phyllaries in 2(–3) series; paleae sometimes surpassing cypselae, margins usually ciliate, rarely eciliate, faces glabrous or hairy; pappi usually coroniform or of 2–6+ scales, sometimes 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis)
→ 2
2. Leaves often bluish green, glaucous; receptacles usually conic to columnar (paleae not surpassing cypselae, except in R. laciniata: paleae surpassing cypselae, receptacles sometimes hemispheric to ovoid, disc corolla lobes yellow); ray florets 0 or 8–15+, corollas bright yellow; disc corollas proximally yellow to yellowish green, distally yellow or greenish to brown-purple; cypselae (3–)3.5–7.5 mm; pappi coroniform, or of 2–6 scales 0.1–2.5 mm
sect. Macrocline
2. Leaves green, not glaucous; receptacles usually conic to hemispheric, rarely columnar (paleae surpassing cypselae); ray florets 6–25+, corollas usually yellow-orange proximally, yellow distally, sometimes with basal maroon splotch (orangish red to maroon in R. graminifolia); disc corollas proximally yellow to yellowish green, distally usually brown-purple, lobes sometimes yellowish or greenish; cypselae 1.5–3.5(–4) mm; pappi coroniform, or of 8+ unequal scales 0.1–2 mm, or 0 (sometimes cypselae each with glandular hairs around apices in R. heliopsidis)
sect. Rudbeckia
Source FNA vol. 21, p. 48. FNA vol. 21, p. 44. Authors: Lowell E. Urbatsch, Patricia B. Cox.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia > sect. Macrocline Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae
Sibling taxa
R. alpicola, R. amplexicaulis, R. auriculata, R. californica, R. fulgida, R. graminifolia, R. grandiflora, R. heliopsidis, R. hirta, R. klamathensis, R. laciniata, R. maxima, R. missouriensis, R. mohrii, R. mollis, R. montana, R. nitida, R. occidentalis, R. scabrifolia, R. subtomentosa, R. texana, R. triloba
Subordinate taxa
R. sect. Dracopis, R. sect. Macrocline, R. sect. Rudbeckia
Synonyms R. californica var. glauca
Name authority Eastwood: Leafl. W. Bot. 2: 55. (1937) Linnaeus: Sp. Pl. 2: 906. (1753): Gen. Pl. ed. 5, 387. (1754)
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