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California coneflower, waxy cone-flower

Habit Perennials, to 150 cm (roots fibrous). Annuals, biennials, or perennials, 20–300 cm.
Leaves

bluish green (heavily glaucous), blades lanceolate to elliptic (not lobed), leathery, bases attenuate, margins entire or remotely serrulate, apices acute, faces glabrous;

basal petiolate, 20–50 × 4–10 cm;

cauline petiolate or sessile, 10–25 × 2–8 cm.

basal and cauline or mostly cauline; alternate; petiolate or sessile;

blades mostly deltate, elliptic, lanceolate, linear, lyrate, oblanceolate, oblong, ovate, pandurate, or spatulate, often 1–2-pinnately lobed or -pinnatifid, ultimate margins entire, dentate, serrate, or toothed, faces glabrous or hairy, often gland-dotted.

Involucres

hemispheric to rotate.

Receptacles

conic to columnar;

paleae 4–6.5 mm, apices acute, often attenuate, abaxial tips hairy.

subhemispheric, ovoid, conic, or columnar (8–70+ mm high), paleate (paleae strongly conduplicate, each partly investing its subtended floret).

Ray florets

7–15;

laminae elliptic to oblong, 25–40 × 8–14 mm, abaxially hairy.

0 or 3–25+, neuter;

corollas yellow or orange, brown-purple, maroon, or reddish, sometimes bicolor (orange, brown-purple, maroon, or reddish plus yellow, laminae often drooping or reflexed).

Disc florets

250–400+;

corollas yellowish green, 3–4 mm;

style branches ca. 1 mm, apices acute.

50–800+, bisexual, fertile;

corollas yellowish green to yellow or brown-purple, tubes shorter than or equaling funnelform or cylindric throats, lobes 5, deltate to triangular or obovate;

anther thecae dark;

stigmatic papillae in 2 lines.

Phyllaries

to 1.5 cm.

persistent, (5–)15–30+ in 1–3 series (distinct, elliptic, lanceolate, linear, ovate, or triangular, usually subequal, sometimes unequal, outer longer, all herbaceous, distally or throughout, or inner scarious, at least outer soon reflexed).

Calyculi

0.

Heads

borne singly or (2–10) in ± corymbiform arrays.

usually radiate, sometimes discoid, borne singly or in loose, ± corymbiform or paniculiform arrays.

Cypselae

4–5.5 mm;

pappi coroniform or of ± connate scales, to 1.2 mm.

subterete, or obpyramidal and 4-angled, or strongly compressed and linear-oblanceolate to oblong-oblique, glabrous or hairy;

pappi 0, or coroniform, or of 2–8+ scales or teeth.

Discs

15–35 × 14–22 mm.

2n

= 36.

Rudbeckia glaucescens

Asteraceae tribe Heliantheae subtribe Rudbeckiinae

Phenology Flowering summer–fall.
Habitat Meadows, seeps, streamsides
Elevation 60–1300 m (200–4300 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
North America; Mexico
Discussion

Rudbeckia glaucescens often grows on serpentine and often with Darlingtonia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 2, species 30 (2 genera, 27 species in the flora).

The genera of Rudbeckiinae have traditionally been included in Verbesininae (e.g., G. Bentham 1873) or in Helianthinae (e.g., T. F. Stuessy 1977[1978]). Sometimes, Rudbeckiinae has included Echinacea (e.g., P. O. Karis and O. Ryding 1994). Here, we follow H. Robinson (1981) in placing Echinacea in Ecliptinae.

In keys and descriptions for Ratibida and Rudbeckia, “discs” refers to receptacles at late flowering with paleae and florets in place and included in assessing shapes and in measurements of lengths and diameters.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres (early flowering) hemispheric to rotate, 15–30+ mm diam.; phyllaries 15–30+ in 2–3 series, subequal; cypselae ± 4-angled, not strongly compressed, margins not pectinate or ciliate
Rudbeckia
1. Involucres rotate, 8–12+ mm diam.; phyllaries 14–28+ in 2 series, unequal (outer notably longer than inner); cypselae strongly compressed, abaxial margin of each usually pectinate or ciliate
Ratibida
Source FNA vol. 21, p. 48. FNA vol. 21, p. 42.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Rudbeckiinae > Rudbeckia > sect. Macrocline Asteraceae > tribe Heliantheae
Sibling taxa
R. alpicola, R. amplexicaulis, R. auriculata, R. californica, R. fulgida, R. graminifolia, R. grandiflora, R. heliopsidis, R. hirta, R. klamathensis, R. laciniata, R. maxima, R. missouriensis, R. mohrii, R. mollis, R. montana, R. nitida, R. occidentalis, R. scabrifolia, R. subtomentosa, R. texana, R. triloba
Subordinate taxa
Ratibida, Rudbeckia
Synonyms R. californica var. glauca
Name authority Eastwood: Leafl. W. Bot. 2: 55. (1937) H. Robinson: Phytologia 41: 43. (1978)
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