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California blackberry, California or Pacific or creeping blackberry, dewberry, Douglasberry, dwarf red raspberry, Pacific blackberry, Pacific dewberry, Pacific trailing blackberry, salmonberry, trailing blackberry

American red raspberry, common red raspberry, framboisier, nagoonberry, North American red raspberry, red raspberry, western red raspberry

Habit Shrubs, usually dioecious, to 2 dm, armed. Shrubs, 5–25 dm, armed or unarmed.
Stems

biennial, creeping (often growing over other vegetation), sparsely to densely hairy, eglandular or sparsely to densely sessile- to stipitate-glandular, usually strongly pruinose;

prickles moderate to dense, erect to retrorse or hooked, weak to moderately stout, 4–10 mm, narrow- to broad-based.

biennial, erect, glabrescent, eglandular or stipitate-glandular, strongly pruinose;

prickles absent or sparse to dense, erect, weak, 1–4 mm, broad-based;

bristles present or absent.

Leaves

deciduous or persistent, usually ternate, sometimes simple or pinnately compound;

stipules filiform to linear, 7–17 mm;

leaflets 3(–5), terminal ovate to lanceolate, 3.5–12 × 3–10 cm, base cuneate or rounded to cordate, shallowly sharp-lobed or unlobed, margins coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces with slender, erect prickles on larger veins, moderately to densely hairy, eglandular, rarely sparsely sessile-glandular along midvein.

deciduous, pinnately compound;

stipules filiform, 5–10 mm;

petiole unarmed or with prickles, strigose or glabrous, glandular or eglandular;

terminal leaflets petiolulate, lateral sessile or subsessile;

leaflets 3–5(–7), terminal ovate to lanceolate, 7–15 × 4–11 cm, base rounded to cordate, unlobed or 2-lobed, margins serrate to doubly serrate, apex acute to attenuate, abaxial surfaces unarmed or with prickles, strongly white-tomentose, eglandular or glandular.

Inflorescences

terminal on short shoots, usually appearing axillary, 1–5-flowered, cymiform or racemiform.

terminal and axillary, (1–)3–7(–20)-flowered, racemiform.

Pedicels

prickles moderate to dense, erect, densely hairy, eglandular or sparsely to densely stipitate-glandular.

unarmed or prickles sparse, hooked, glabrous or sparsely pubescent, stipitate-glandular or eglandular.

Flowers

usually functionally unisexual;

petals white, in staminate flowers ovate or obovate to narrowly elliptic, 7–18 mm, in pistillate ovate to elliptic, 6–11 mm;

filaments filiform;

ovaries glabrous or hairy.

bisexual;

petals white to greenish white, spatulate to obovate, 5–10 mm;

filaments filiform or slightly dilated basally;

ovaries pubescent.

Fruits

black, sometimes red or purple, rarely white, not pruinose, globose to cylindric, 1–2.5 cm;

drupelets 20–50, strongly coherent, separating with torus attached.

usually red to whitish, rarely amber, globose to conic, 0.5–2 cm;

drupelets 10–60, coherent, separating from torus.

2n

= 42, 56, 63, 70, 77, 84, 91.

Rubus ursinus

Rubus idaeus

Phenology Flowering (Jan–)Mar–Aug(–Sep).
Habitat Woodlands, shrublands, open or disturbed areas, dry to damp soil
Elevation 0–1600 m (0–5200 ft)
Distribution
from FNA
CA; ID; MT; OR; WA; BC; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; NU; ON; PE; QC; SK; YT; Mexico; Eurasia; Africa
[WildflowerSearch map]
[BONAP county map]
Discussion

In habit, Rubus ursinus is often reminiscent of the predominantly eastern North American, non-sympatric R. flagellaris; it is also polymorphic. Rubus ursinus comprises a polyploid spectrum dominated by octoploid and dodecaploid plants. It is an allopolyploid involving phylogenetically distant ancestors with its closest relative being the Hawaiian endemic R. macraei A. Gray. See S. W. Brown (1943) for a discussion of morphological and chromosome number variation in R. ursinus. The widespread, glaucous-stemmed, trifoliate-leaved forms of R. ursinus are vegetatively nearly identical to R. caesius. Of agricultural significance, R. ursinus is a parent of some important cultivars, including loganberry and boysenberry.

There has been uncertainty whether the name Rubus menziesii Hooker and subsequent combinations [Parmena menziesii (Hooker) Greene, R. spectabilis var. menziesii (Hooker) S. Watson, and R. ursinus var. menziesii (Hooker) Focke] pertain to R. spectabilis or R. ursinus. The protologue by Hooker describes a hairy, relatively small, procumbent plant, which is congruent with R. ursinus. The type specimen at Kew also looks like R. ursinus. In the protologue Hooker also claimed the species has red petals and suggested that it may have an affinity with R. spectabilis.

Specimens examined for this study of North American collections attributed to Rubus macrophyllus Weihe & Nees (synonym R. amplificatus Lees) are R. ursinus. If R. macrophyllus was once present in the flora area it is likely absent now; some reports of R. macrophyllus could represent hybrids. There are reports of R. ursinus hybridizing with R. bifrons and R. pensilvanicus in California (T. S. Mallah 1954; L. V. Clark and M. Jasieniuk 2012; L. A. Alice, unpubl.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Rubus idaeus is the source of most of the cultivated red and amber raspberries. Plants were used by the Cherokee, Cree, and others as a parturient; an infusion of dried leaves in water was used to prepare the uterus for childbirth and help women recover after childbirth (D. E. Moerman 1998; D. Hoffman 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems eglandular 13a
subsp. idaeus
1. Stems stipitate-glandular.
subsp. strigosus
Source FNA vol. 9, p. 55. FNA vol. 9, p. 42.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Subordinate taxa
R. idaeus subsp. idaeus, R. idaeus subsp. strigosus
Synonyms R. eastwoodianus, R. macropetalus, R. sirbenus, R. ursinus var. eastwoodianus, R. ursinus subsp. macropetalus, R. ursinus var. macropetalus, R. ursinus var. sirbenus, R. vitifolius, R. vitifolius var. eastwoodianus, R. vitifolius subsp. ursinus
Name authority Chamisso & Schlechtendal: Linnaea 2: 11. (1827) Linnaeus: Sp. Pl. 1: 492. (1753)
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