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elm-leaf blackberry

black raspberry, blackcap, blackcap raspberry, dark raspberry, dwarf bramble, western black raspberry, white-bark raspberry

Habit Shrubs, 30–50 dm, unarmed or armed. Shrubs, 10–30 dm, armed.
Stems

biennial, erect, then arching, usually moderately, rarely sparsely, hairy hairs stellate, eglandular, strongly pruinose;

prickles absent or sparse to dense, erect to hooked, stout, 3–10 mm, broad-based.

biennial, ascending, often arching, glabrous or sparsely puberulent, eglandular or sparsely to densely short-stipitate-glandular, strongly pruinose;

prickles moderate to dense, erect or curved, slender to stout, 4–8 mm, usually broad-based, sometimes narrow-based.

Leaves

persistent or nearly so, ternate to palmately compound;

stipules filiform to linear, 4–12 mm;

leaflets 3–5, terminal ovate, elliptic, or suborbiculate to obovate, 4–10 × 2–6(–9) cm, base rounded to cuneate, unlobed, margins finely to moderately serrate, apex acute or acuminate to attenuate, abaxial surfaces with hooked prickles on larger veins, densely white short-velutinous, hairs usually minute, eglandular.

deciduous, palmately compound or ternate;

stipules filiform, 5–10 mm;

petiole with hooked or erect prickles, glabrous, eglandular or sparsely to densely sessile- to short-stipitate-glandular;

lateral leaflets stalked, at least 2 larger ones, often with prickles, glabrous, eglandular or stipitate-glandular;

leaflets 3–5(–7), terminal ovate to lanceolate, 6–10 × 3–6 cm, base cordate to truncate, unlobed or shallowly 3-lobed, margins serrate to doubly serrate, apex acuminate, abaxial surface sometimes with prickles on midrib and some lateral veins, glabrous to strongly white-tomentose, usually eglandular.

Inflorescences

terminal, 10–60-flowered, paniculiform or thyrsiform.

axillary and terminal, (1–)3–10(–12)-flowered, flat-topped cymiform or umbelliform.

Pedicels

unarmed or prickles moderate to dense, erect to hooked, densely short-hairy, eglandular.

prickles sparse to dense, hooked or erect, puberulent, eglandular or sparsely to densely sessile- or short-stipitate-glandular.

Flowers

bisexual;

petals usually pink, sometimes white, obovate or elliptic to suborbiculate, 5–12 mm;

filaments filiform;

ovaries apically hairy.

bisexual;

petals erect, then ascending, white, oblong to oblanceolate, 3–6(–8) mm;

filaments laminar;

ovaries pubescent, rarely glabrous.

Fruits

black, globose to ovoid, 1–1.5 cm;

drupelets 10–40, strongly coherent, separating with torus attached.

reddish purple to nearly black, depressed globose to conic, 1–2 cm;

drupelets 20–60, coherent, separating from torus.

2n

= 14.

= 14.

Rubus ulmifolius

Rubus leucodermis

Phenology Flowering May–Jul.
Habitat Woodland edges, open areas, disturbed areas, moist soil
Elevation 0–500 m (0–1600 ft)
Distribution
from FNA
CA; NJ; NV; OR; WA; s Europe; c Europe; n Africa [Introduced in North America; introduced also in s South America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Rubus ulmifolius can be distinguished, especially from the closely related R. bifrons and R. vestitus, by its strongly pruinose stems, finely serrate leaflets, and lack of glands throughout. Unlike R. ulmifolius, strongly pruinose native Rubus species lack relatively large and showy pink petals. Some new stems developing from tip-rooting, and early leaves on such stems (especially in shade), are not whitened abaxially and are tomentose. Such unusual stems develop typical leaves and surfaces in parts formed later.

There has been nomenclatural confusion over unarmed plants of Rubus ulmifolius (see E. Monasterio-Huelin and H. E. Weber 1996). Rubus inermis Willdenow is an illegitimate later homonym of R. inermis Pourret, and the type specimens for both names are of poor-quality. If an unarmed variety of R. ulmifolius is to be recognized, such as occurs in California, the name var. anoplothyrsus Sudre should be used instead of var. inermis (Willdenow) Focke.

Rubus ulmifolius is attractive and is potentially weedy but relatively uncommon, found primarily in coastal California from the San Francisco Bay region southward. It probably no longer persists in New Jersey, last collected there in 1897, and it is rare in Nevada and Oregon. Rubus ulmifolius likely hybridizes with R. bifrons where they co-occur; it is one of relatively few diploid and sexually reproducing species of European blackberries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Rubus leucodermis is similar to R. occidentalis, differing primarily by its leaflets having acuminate apices and lateral leaflets that are distinctly stalked. Prickles on the pedicels of R. occidentalis are erect; those of R. leucodermis tend to be hooked.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves abaxially white-tomentose; drupelets (20–)30–60.
var. leucodermis
1. Leaves abaxially glabrous, rarely sparsely hairy; drupelets 20–40.
var. nigerrimus
Source FNA vol. 9, p. 54. FNA vol. 9, p. 44.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Subordinate taxa
R. leucodermis var. leucodermis, R. leucodermis var. nigerrimus
Synonyms R. discolor, R. ulmifolius var. anoplothyrsus R. occidentalis subsp. leucodermis
Name authority Schott: Isis (Oken) 1818: 821. (1818) Douglas ex Torrey & A. Gray: Fl. N. Amer. 1: 454. (1840)
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