Rubus ulmifolius |
Rubus lasiococcus |
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elm-leaf blackberry |
dwarf bramble, hairy-fruit smooth dewberry, rough fruit berry, roughfruit raspberry |
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Habit | Shrubs, 30–50 dm, unarmed or armed. | Herbs, 1–1.5 dm, unarmed. |
Stems | biennial, erect, then arching, usually moderately, rarely sparsely, hairy hairs stellate, eglandular, strongly pruinose; prickles absent or sparse to dense, erect to hooked, stout, 3–10 mm, broad-based. |
creeping, flowering branches erect, sparsely hairy, eglandular or sparsely to moderately short-stipitate-glandular, not pruinose. |
Leaves | persistent or nearly so, ternate to palmately compound; stipules filiform to linear, 4–12 mm; leaflets 3–5, terminal ovate, elliptic, or suborbiculate to obovate, 4–10 × 2–6(–9) cm, base rounded to cuneate, unlobed, margins finely to moderately serrate, apex acute or acuminate to attenuate, abaxial surfaces with hooked prickles on larger veins, densely white short-velutinous, hairs usually minute, eglandular. |
deciduous, simple or ternate; stipules broadly elliptic to ovate, 5–10 mm; leaflets 3(–5), blade reniform or orbiculate, terminal lobe or leaflets broadly obovate to obovate-cuneate, 1.5–4(–6) × 1.5–4(–6) cm, base cordate, 3(–5)-lobed, margins coarsely doubly dentate, apex rounded to acute, abaxial surfaces sparsely hairy on veins, eglandular. |
Inflorescences | terminal, 10–60-flowered, paniculiform or thyrsiform. |
1–2-flowered. |
Pedicels | unarmed or prickles moderate to dense, erect to hooked, densely short-hairy, eglandular. |
sparsely to moderately hairy, short-stipitate-glandular. |
Flowers | bisexual; petals usually pink, sometimes white, obovate or elliptic to suborbiculate, 5–12 mm; filaments filiform; ovaries apically hairy. |
bisexual; petals white, broadly elliptic to obovate to orbiculate, (5–)8–10(–12) mm; filaments filiform; ovaries densely hairy, styles filiform, glabrous. |
Fruits | black, globose to ovoid, 1–1.5 cm; drupelets 10–40, strongly coherent, separating with torus attached. |
red, hemispheric, 1 cm; drupelets 5–15, weakly coherent, falling separately or as loose units free from torus. |
2n | = 14. |
= 14. |
Rubus ulmifolius |
Rubus lasiococcus |
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Phenology | Flowering May–Jul. | Flowering May–Aug. |
Habitat | Woodland edges, open areas, disturbed areas, moist soil | Moist to dry, semiopen forests, wet meadows, roadsides, dry sand |
Elevation | 0–500 m (0–1600 ft) | 800–2000 m (2600–6600 ft) |
Distribution |
CA; NJ; NV; OR; WA; s Europe; c Europe; n Africa [Introduced in North America; introduced also in s South America]
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CA; OR; WA; BC
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Discussion | Rubus ulmifolius can be distinguished, especially from the closely related R. bifrons and R. vestitus, by its strongly pruinose stems, finely serrate leaflets, and lack of glands throughout. Unlike R. ulmifolius, strongly pruinose native Rubus species lack relatively large and showy pink petals. Some new stems developing from tip-rooting, and early leaves on such stems (especially in shade), are not whitened abaxially and are tomentose. Such unusual stems develop typical leaves and surfaces in parts formed later. There has been nomenclatural confusion over unarmed plants of Rubus ulmifolius (see E. Monasterio-Huelin and H. E. Weber 1996). Rubus inermis Willdenow is an illegitimate later homonym of R. inermis Pourret, and the type specimens for both names are of poor-quality. If an unarmed variety of R. ulmifolius is to be recognized, such as occurs in California, the name var. anoplothyrsus Sudre should be used instead of var. inermis (Willdenow) Focke. Rubus ulmifolius is attractive and is potentially weedy but relatively uncommon, found primarily in coastal California from the San Francisco Bay region southward. It probably no longer persists in New Jersey, last collected there in 1897, and it is rare in Nevada and Oregon. Rubus ulmifolius likely hybridizes with R. bifrons where they co-occur; it is one of relatively few diploid and sexually reproducing species of European blackberries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Rubus lasiococcus is recognized by its creeping, unarmed stems, simple 3-lobed to 3-foliate leaves, small flowers, white petals, and densely hairy ovaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 54. | FNA vol. 9, p. 44. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus |
Sibling taxa | ||
Synonyms | R. discolor, R. ulmifolius var. anoplothyrsus | |
Name authority | Schott: Isis (Oken) 1818: 821. (1818) | A. Gray: Proc. Amer. Acad. Arts 17: 201. (1882) |
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