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southern dewberry

European blackberry

Habit Shrubs, to 3(–7) dm, sometimes climbing higher through other vegetation, moderately to densely armed. Shrubs, 10–20 dm, armed.
Stems

biennial, initially low-arching, then falling and creeping (or climbing through other vegetation), glabrous or moderately hairy, sparsely to densely short- to long-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved, sometimes distally slender, 1–4 mm, broad-based;

bristles absent or sparse to dense, erect to retrorse, red to purple, rarely green, slender, weak, gland-tipped.

biennial, arching, usually densely, sometimes moderately, hairy, usually eglandular, sometimes sessile- to short-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved to erect, slender, 3–10 mm, broad-based.

Leaves

persistent or semipersistent, ternate to palmately compound, lustrous;

stipules filiform, linear, or lanceolate, 2–12(–15) mm;

leaflets 3–5, terminal narrowly elliptic or ovate to obovate, 2–8.5 × 0.7–4.5 cm, base rounded to cuneate, unlobed, margins moderately to coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces with hooked prickles on midvein, glabrous or sparsely to moderately hairy, eglandular or sparsely short-stipitate-glandular along central vein.

deciduous or ± persistent, palmately compound;

stipules filiform to linear, 5–15 mm;

leaflets 3–5, terminal usually suborbiculate to orbiculate, sometimes broadly elliptic, 4–9 × 4–8 cm, base rounded to shallowly cordate, unlobed, margins moderately to coarsely serrate, apex acute to cuspidate, abaxial surfaces with hooked prickles on midvein, densely white-gray to gray-green tomentose, eglandular or obscurely short-stipitate- to sessile-glandular along midveins.

Inflorescences

terminal on short shoots, usually appearing axillary, 1(–3)-flowered.

terminal, 7–35-flowered, cymiform or thyrsiform.

Pedicels

prickles and, often, bristles moderate to dense, recurved, moderately to densely hairy, sparsely to moderately sessile- to short-stipitate-glandular.

prickles moderate to dense, recurved to erect, densely hairy, moderately to densely stipitate-glandular.

Flowers

bisexual;

petals white to pink, elliptic to obovate, 10–16(–25) mm;

filaments filiform;

ovaries glabrous.

bisexual;

petals pink to magenta, elliptic to suborbiculate, 6–12(–15) mm;

filaments filiform;

ovaries apically hairy.

Fruits

black, globose to ovoid, 1–1.5(–2) cm;

drupelets 10–50, strongly coherent, separating with torus attached.

black, globose, 1–1.5 cm;

drupelets 15–40, coherent, separating with torus attached.

2n

= 14.

= 28.

Rubus trivialis

Rubus vestitus

Phenology Flowering Jan–Jun. Flowering Jul–Sep.
Habitat Open woodlands, savannas, prairies, meadows, sand dunes, disturbed areas, dry to seasonally wet soil Woodland edges, open areas, disturbed areas, in moist soil
Elevation 0–200 m (0–700 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
AL; AR; FL; GA; IL; KS; KY; LA; MO; MS; NC; OK; SC; TN; TX; VA; Mexico (Coahuila, Nuevo León, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; BC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Rubus trivialis is distinguished from other species of Rubus by its frequently glandular-bristly and generally creeping stems, abundant recurved prickles, and typically persistent or semipersistent, lustrous primocane leaves with relatively narrow leaflets. Although emerging primocanes typically reach to 30 cm above the ground, vigorous plants can have new primocanes standing erect to 70 cm that later fall to the ground or onto adjacent vegetation as they continue to enlarge. L. H. Bailey (1941–1945) believed that there was no clear separation between members of sect. Persistentes Fernald (referred to by Bailey as sect. Verotriviales [illegitimate sectional name for R. trivialis and associated species]) and sect. Procumbentes (by Bailey as sect. Flagellares; including R. flagellaris and related species). All characteristics that he recognized for distinguishing sect. Persistentes from sect. Procumbentes are often present in the latter, with the exception of hispid-setose stems, and glabrous plants of R. trivialis are difficult to distinguish from some plants of R. flagellaris; to include the latter species within the former would be reasonable.

Rubus ×inferior L. H. Bailey is a putative hybrid of R. trivialis and R. cuneifolius.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus vestitus can be distinguished, especially from the closely related R. bifrons and R. ulmifolius, by its long-stipitate-glandular hairs, particularly in the inflorescence, and terminal primocane leaflets that are typically suborbiculate and abaxially densely tomentose. Rubus allegheniensis can possess similar glandular hairs; it does not have pink petals, thyrsiform inflorescences, and rounded and abaxially densely tomentose, often whitened terminal leaflets.

Rubus vestitus has not yet been found in Washington, but it is likely there. Because of the overall similarity of R. vestitus to R. bifrons, it is possible that this potentially weedy species is widespread, but rare and possibly overlooked, occurring as far south as California. In Oregon, R. vestitus prefers shady habitats (B. Wilson, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 54. FNA vol. 9, p. 56.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus
Synonyms R. agilis, R. continentalis, R. duplaris, R. ictus, R. lucidus, R. macvaughii, R. mirus, R. nessianus, R. riograndis, R. rubrisetus, R. sons, R. tallahasseanus
Name authority Michaux: Fl. Bor.-Amer. 1: 296. (1803) Weihe & Nees: in M. J. Bluff et al., Comp. Fl. German. 1: 684. (1825)
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