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southern dewberry

stone bramble

Habit Shrubs, to 3(–7) dm, sometimes climbing higher through other vegetation, moderately to densely armed. Herbs or subshrubs, to 4 dm, armed or unarmed.
Stems

biennial, initially low-arching, then falling and creeping (or climbing through other vegetation), glabrous or moderately hairy, sparsely to densely short- to long-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved, sometimes distally slender, 1–4 mm, broad-based;

bristles absent or sparse to dense, erect to retrorse, red to purple, rarely green, slender, weak, gland-tipped.

usually creeping, ± woody basally, sparsely to densely hairy, eglandular, not pruinose;

prickles absent or sparse, erect, weak, 1–3 mm, narrow-based.

Leaves

persistent or semipersistent, ternate to palmately compound, lustrous;

stipules filiform, linear, or lanceolate, 2–12(–15) mm;

leaflets 3–5, terminal narrowly elliptic or ovate to obovate, 2–8.5 × 0.7–4.5 cm, base rounded to cuneate, unlobed, margins moderately to coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces with hooked prickles on midvein, glabrous or sparsely to moderately hairy, eglandular or sparsely short-stipitate-glandular along central vein.

deciduous, ternate;

stipules free from petioles, lanceolate to ovate or elliptic, 6–11 mm;

petiole 2–11 cm;

petiolule of terminal leaflet 8–20 mm;

lateral leaflet sessile or subsessile;

terminal leaflets ovate to elliptic, 4–8 × 3–7 cm, base tapered, unlobed, lateral leaflets sometimes shallowly lobed, margins coarsely doubly serrate, apex acute, abaxial surfaces unarmed, sparsely hairy, eglandular.

Inflorescences

terminal on short shoots, usually appearing axillary, 1(–3)-flowered.

axillary, 1–6-flowered, cymiform or umbelliform.

Pedicels

prickles and, often, bristles moderate to dense, recurved, moderately to densely hairy, sparsely to moderately sessile- to short-stipitate-glandular.

unarmed or prickles weak, erect, hairy, eglandular or sparsely stipitate-glandular.

Flowers

bisexual;

petals white to pink, elliptic to obovate, 10–16(–25) mm;

filaments filiform;

ovaries glabrous.

bisexual;

petals greenish white, narrowly obovate, 5.5–7 mm;

filaments laminar;

ovaries glabrous.

Fruits

black, globose to ovoid, 1–1.5(–2) cm;

drupelets 10–50, strongly coherent, separating with torus attached.

red, globose, 0.5–1.5 cm;

drupelets 1–10, not or weakly coherent, separating with torus attached.

2n

= 14.

= 28.

Rubus trivialis

Rubus saxatilis

Phenology Flowering Jan–Jun. Flowering May–Jul.
Habitat Open woodlands, savannas, prairies, meadows, sand dunes, disturbed areas, dry to seasonally wet soil Birch scrub, talus slopes
Elevation 0–200 m (0–700 ft) 1300–2100 m (4300–6900 ft)
Distribution
from FNA
AL; AR; FL; GA; IL; KS; KY; LA; MO; MS; NC; OK; SC; TN; TX; VA; Mexico (Coahuila, Nuevo León, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from USDA
Greenland; Eurasia
[BONAP county map]
Discussion

Rubus trivialis is distinguished from other species of Rubus by its frequently glandular-bristly and generally creeping stems, abundant recurved prickles, and typically persistent or semipersistent, lustrous primocane leaves with relatively narrow leaflets. Although emerging primocanes typically reach to 30 cm above the ground, vigorous plants can have new primocanes standing erect to 70 cm that later fall to the ground or onto adjacent vegetation as they continue to enlarge. L. H. Bailey (1941–1945) believed that there was no clear separation between members of sect. Persistentes Fernald (referred to by Bailey as sect. Verotriviales [illegitimate sectional name for R. trivialis and associated species]) and sect. Procumbentes (by Bailey as sect. Flagellares; including R. flagellaris and related species). All characteristics that he recognized for distinguishing sect. Persistentes from sect. Procumbentes are often present in the latter, with the exception of hispid-setose stems, and glabrous plants of R. trivialis are difficult to distinguish from some plants of R. flagellaris; to include the latter species within the former would be reasonable.

Rubus ×inferior L. H. Bailey is a putative hybrid of R. trivialis and R. cuneifolius.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus saxatilis is defined by its creeping, unarmed or weakly armed stems, ternate leaves, and relatively small white petals. It has very weak prickles that detach easily, sometimes giving plants an unarmed appearance. The species is known in the flora area from only two sites in southern Greenland (T. W. Böcher 1938; D. B. Jensen and K. D. Christensen 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 54. FNA vol. 9, p. 52.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Synonyms R. agilis, R. continentalis, R. duplaris, R. ictus, R. lucidus, R. macvaughii, R. mirus, R. nessianus, R. riograndis, R. rubrisetus, R. sons, R. tallahasseanus
Name authority Michaux: Fl. Bor.-Amer. 1: 296. (1803) Linnaeus: Sp. Pl. 1: 494. (1753)
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