Flora of North America species comparison

biennial, initially low-arching, then falling and creeping (or climbing through other vegetation), glabrous or moderately hairy, sparsely to densely short- to long-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved, sometimes distally slender, 1–4 mm, broad-based;

bristles absent or sparse to dense, erect to retrorse, red to purple, rarely green, slender, weak, gland-tipped.

biennial, erect, primocanes and floricanes later over-arching, glabrous or sparsely puberulent, eglandular, strongly pruinose;

prickles sometimes sparse, erect or hooked, narrow to stout, 4–8 mm, narrow- to broad-based.

persistent or semipersistent, ternate to palmately compound, lustrous;

stipules filiform, linear, or lanceolate, 2–12(–15) mm;

leaflets 3–5, terminal narrowly elliptic or ovate to obovate, 2–8.5 × 0.7–4.5 cm, base rounded to cuneate, unlobed, margins moderately to coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces with hooked prickles on midvein, glabrous or sparsely to moderately hairy, eglandular or sparsely short-stipitate-glandular along central vein.

deciduous, ternate or palmately compound;

stipules filiform, 5–10 mm;

petiole usually armed with prickles;

petiolules of terminal leaflets also sometimes armed;

lateral leaflets sessile or subsessile;

leaflets 3(–5), terminal ovate to lanceolate, 5–14 × 3–11 cm, base rounded to cordate, unlobed or laterals sometimes lobed, margins finely to doubly serrate, apex acute, abaxial surfaces sometimes with hooked prickles on midveins, densely white-canescent to tomentose, eglandular.

terminal on short shoots, usually appearing axillary, 1(–3)-flowered.

(2–)3–7(–20)-flowered, cymiform or umbelliform.

prickles and, often, bristles moderate to dense, recurved, moderately to densely hairy, sparsely to moderately sessile- to short-stipitate-glandular.

prickles erect, hooked, puberulent to pubescent, eglandular.

bisexual;

petals white to pink, elliptic to obovate, 10–16(–25) mm;

filaments filiform;

ovaries glabrous.

bisexual;

petals initially erect, later ascending, white, narrowly obovate to elliptic, 5–10 mm, apex sometimes emarginate;

filaments laminar;

ovaries tomentose.

black, globose to ovoid, 1–1.5(–2) cm;

drupelets 10–50, strongly coherent, separating with torus attached.

usually dark purplish to black, rarely amber, pruinose, aromatic, depressed-globose, 1–1.5 cm;

drupelets 20–50, coherent, separating from torus.

= 14.

= 14.