Flora of North America species comparison

biennial, initially low-arching, then falling and creeping (or climbing through other vegetation), glabrous or moderately hairy, sparsely to densely short- to long-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved, sometimes distally slender, 1–4 mm, broad-based;

bristles absent or sparse to dense, erect to retrorse, red to purple, rarely green, slender, weak, gland-tipped.

biennial, ascending, often arching, glabrous or sparsely puberulent, eglandular or sparsely to densely short-stipitate-glandular, strongly pruinose;

prickles moderate to dense, erect or curved, slender to stout, 4–8 mm, usually broad-based, sometimes narrow-based.

persistent or semipersistent, ternate to palmately compound, lustrous;

stipules filiform, linear, or lanceolate, 2–12(–15) mm;

leaflets 3–5, terminal narrowly elliptic or ovate to obovate, 2–8.5 × 0.7–4.5 cm, base rounded to cuneate, unlobed, margins moderately to coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces with hooked prickles on midvein, glabrous or sparsely to moderately hairy, eglandular or sparsely short-stipitate-glandular along central vein.

deciduous, palmately compound or ternate;

stipules filiform, 5–10 mm;

petiole with hooked or erect prickles, glabrous, eglandular or sparsely to densely sessile- to short-stipitate-glandular;

lateral leaflets stalked, at least 2 larger ones, often with prickles, glabrous, eglandular or stipitate-glandular;

leaflets 3–5(–7), terminal ovate to lanceolate, 6–10 × 3–6 cm, base cordate to truncate, unlobed or shallowly 3-lobed, margins serrate to doubly serrate, apex acuminate, abaxial surface sometimes with prickles on midrib and some lateral veins, glabrous to strongly white-tomentose, usually eglandular.

terminal on short shoots, usually appearing axillary, 1(–3)-flowered.

axillary and terminal, (1–)3–10(–12)-flowered, flat-topped cymiform or umbelliform.

prickles and, often, bristles moderate to dense, recurved, moderately to densely hairy, sparsely to moderately sessile- to short-stipitate-glandular.

prickles sparse to dense, hooked or erect, puberulent, eglandular or sparsely to densely sessile- or short-stipitate-glandular.

bisexual;

petals white to pink, elliptic to obovate, 10–16(–25) mm;

filaments filiform;

ovaries glabrous.

bisexual;

petals erect, then ascending, white, oblong to oblanceolate, 3–6(–8) mm;

filaments laminar;

ovaries pubescent, rarely glabrous.

black, globose to ovoid, 1–1.5(–2) cm;

drupelets 10–50, strongly coherent, separating with torus attached.

reddish purple to nearly black, depressed globose to conic, 1–2 cm;

drupelets 20–60, coherent, separating from torus.

= 14.

= 14.