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salmon berry

Chesapeake blackberry, Nanticoke blackberry, topsy

Habit Shrubs, 10–40 dm, usually armed. Shrubs, 7–30 dm, armed.
Stems

erect to arching, glabrate or sparsely to densely hairy, eglandular or sparsely short-stipitate-glandular, rarely densely long-stipitate-glandular, not pruinose;

bark usually papery with age, peeling (especially toward base);

prickles absent or sparse to dense, erect, slender, 1–5 mm, broad- to narrow-based.

biennial, arching, sparsely to densely hairy, sparsely to densely sessile-glandular, rarely short-stipitate-glandular, not pruinose;

bark not papery, peeling;

prickles moderate to dense, hooked, retrorse, or erect, stout, 4–10 mm, broad- or narrow-based.

Leaves

deciduous, ternate;

stipules filiform to linear, 3–10 mm;

terminal leaflets ovate, 4–15 × 3.5–15 cm, base truncate, rounded to shallowly cordate, shallowly, sharply lobed, margins coarsely serrate to doubly serrate, apex acute to acuminate, abaxial surfaces unarmed or with erect prickles on midvein, moderately to densely hairy, eglandular, rarely stipitate-glandular along midvein.

deciduous to semievergreen, palmately compound, rarely ternate;

stipules filiform to linear, rarely narrowly lanceolate, 5–16(–20) mm;

leaflets 3–5, terminal broadly elliptic or ovate, (3.5–)4.5–8.5 × 2.5–6 cm, base rounded, unlobed, margins moderately to coarsely doubly, rarely singly, serrate, apex acute or short-acuminate, abaxial surfaces with hooked prickles on larger veins, short-velutinous to tomentose, sparsely to densely sessile-glandular, rarely short-stipitate-glandular.

Inflorescences

terminal and axillary, 1–2-flowered.

terminal and axillary, 3–15(–25)-flowered, cymiform to thyrsiform, not projected well beyond subtending leaves.

Pedicels

unarmed or prickles sparse, erect, moderately to densely hairy, eglandular, rarely short-stipitate-glandular.

prickles moderate to dense, hooked, retrorse, or erect, densely hairy, eglandular or sparsely to moderately sessile-glandular.

Flowers

bisexual;

petals pink to magenta, broadly to narrowly obovate, 10–30 mm;

filaments laminar;

ovaries glabrous.

bisexual;

petals white to pale pink, obovate or elliptic to orbiculate, (8–)10–15 mm;

filaments filiform;

ovaries glabrous or apically hairy.

Fruits

yellow, orange, or red, globose to ovoid, 1–2 cm;

drupelets 20–80, strongly coherent, separating from torus.

black, globose to subcylindric, 1–2 cm;

drupelets (15–)20–40(–50), strongly coherent, separating with torus attached.

2n

= 14.

= 21, 28.

Rubus spectabilis

Rubus pascuus

Phenology Flowering (Feb–)Mar–Jul. Flowering (Apr–)May–Jun.
Habitat Woodlands, woodland edges, bogs, shorelines, roadsides, disturbed areas, moist to wet soil Woodland edges, swamp margins, dry thickets, open and often disturbed areas, roadsides
Elevation 0–2000 m (0–6600 ft) 0–300(–700) m (0–1000(–2300) ft)
Distribution
from FNA
AK; CA; ID; OR; WA; BC; e Asia (Japan) [Introduced in Europe]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DE; GA; IL; IN; KY; MD; MO; MS; NC; NJ; OK; PA; SC; TN; TX; VA
[BONAP county map]
Discussion

Rubus spectabilis is a thicket-forming shrub that has relatively large and desirably edible fruit. The species is used as an ornamental primarily for its robust, showy flowers and is naturalized in parts of western Europe. It is sister to the Hawaiian endemic R. hawaiiensis A. Gray. See discussion under 36. R. ursinus for the uncertain application of the name R. menziesii Hooker.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus pascuus is probably more widespread than indicated here. The species is most likely to be mistaken for R. bifrons, R. cuneifolius, or R. pensilvanicus. Unlike R. bifrons, R. pascuus is generally more densely glandular, has much shorter, compact inflorescences that usually barely surpass the leaves, and tends to hold its primocane leaves relatively erect (versus spreading in R. bifrons). Although both can be semievergreen, R. pascuus will lose its leaves more readily than R. bifrons in areas where they co-occur. When compared with R. cuneifolius, R. pascuus has pale pink to white petals and ovate to broadly elliptic leaflets with flat margins, the entire part restricted to the base; R. cuneifolius has white petals and cuneate to obovate leaflets with usually revolute margins, the proximal third entire. When compared with R. pensilvanicus, R. pascuus has pale pink to white petals. Rubus pensilvanicus only has white petals as well as abaxial leaf surfaces, which are white to gray-green in R. pascuus with dense, short-velutinous to tomentose indument, while those of R. pensilvanicus can be hairy but never whitish to gray-green. A notable aspect of R. pascuus is how vigorously armed it is, more so than most other Rubus species in the flora area.

Rubus pascuus may represent a stabilized hybrid between a European and a native blackberry species, suggested by whitish pubescence on the abaxial leaf surface and pale pink petals. L. H. Bailey placed R. pascuus in his sect. Cuneifolii (L. H. Bailey) L. H. Bailey, likely because of the whitish hairs on the abaxial leaf surface.

The use of the name Rubus pascuus here is tentative; it is the best fit to the material observed. This treatment follows H. A. Davis (1990) for the choice of the name to use and its associated synonymy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 53. FNA vol. 9, p. 49.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Synonyms R. franciscanus, R. spectabilis var. franciscanus
Name authority Pursh: Fl. Amer. Sept. 1: 348, plate 16. (1813) L. H. Bailey: Gentes Herb. 5: 440, fig. 200. (1943)
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