Rubus setosus |
Rubus saxatilis |
|
---|---|---|
bristly blackberry, ronce sétuleuse, setose blackberry |
stone bramble |
|
Habit | Shrubs, 2–10(–15) dm, armed. | Herbs or subshrubs, to 4 dm, armed or unarmed. |
Stems | biennial, erect to arching, rarely creeping, not node- or tip-rooting, sparsely to moderately hairy, sparsely to densely long-, rarely short-stipitate-glandular, glands rounded to flattened, not pruinose; prickles absent or sparsely to moderately dense, erect to retrorse, weak, slender, 1–3(–5) mm, narrow-based; bristles sparse to dense, erect to retrorse, green to reddish, narrow, semirigid, sometimes flexible, not gland-tipped. |
usually creeping, ± woody basally, sparsely to densely hairy, eglandular, not pruinose; prickles absent or sparse, erect, weak, 1–3 mm, narrow-based. |
Leaves | deciduous, ternate or palmately compound, sometimes ± lustrous; stipules filiform to lanceolate, 5–20(–38) mm; leaflets (3–)5, terminal obovate or elliptic to rhombic, 4.5–11.5 × 2–7 cm, base cuneate to narrowly rounded, unlobed, margins moderately, coarsely serrate to doubly serrate or serrate-dentate, apex acute or acuminate to short-attenuate, abaxial surfaces unarmed or with bristles on midvein, sparsely to moderately hairy, eglandular or sparsely to moderately sessile- to long-stipitate-glandular along larger veins. |
deciduous, ternate; stipules free from petioles, lanceolate to ovate or elliptic, 6–11 mm; petiole 2–11 cm; petiolule of terminal leaflet 8–20 mm; lateral leaflet sessile or subsessile; terminal leaflets ovate to elliptic, 4–8 × 3–7 cm, base tapered, unlobed, lateral leaflets sometimes shallowly lobed, margins coarsely doubly serrate, apex acute, abaxial surfaces unarmed, sparsely hairy, eglandular. |
Inflorescences | terminal on short shoots, sometimes appearing axillary, 5–15(–20)-flowered, racemiform, cymiform, or thyrsiform. |
axillary, 1–6-flowered, cymiform or umbelliform. |
Pedicels | prickles or bristles sparse to dense, erect to retrorse, moderately to densely hairy, moderately to densely stipitate-glandular. |
unarmed or prickles weak, erect, hairy, eglandular or sparsely stipitate-glandular. |
Flowers | bisexual; petals white, obovate to oblanceolate, 6–13 mm; filaments filiform; ovaries glabrous. |
bisexual; petals greenish white, narrowly obovate, 5.5–7 mm; filaments laminar; ovaries glabrous. |
Fruits | black, globose, 0.7–1.5 cm; drupelets 5–25, strongly coherent, separating with torus attached. |
red, globose, 0.5–1.5 cm; drupelets 1–10, not or weakly coherent, separating with torus attached. |
2n | = 14, 21, 28, 35. |
= 28. |
Rubus setosus |
Rubus saxatilis |
|
Phenology | Flowering Jun–Jul(–Aug). | Flowering May–Jul. |
Habitat | Open woodlands, savannas, prairies, meadows, disturbed areas, dry to wet soil | Birch scrub, talus slopes |
Elevation | 0–1000 m (0–3300 ft) | 1300–2100 m (4300–6900 ft) |
Distribution |
CT; DC; DE; IA; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; NB; NF; NS; ON; PE; QC
|
Greenland; Eurasia |
Discussion | Rubus setosus is identified by erect to arching, rarely creeping, not tip-rooting, terete primocanes, always setose but variable in density, and deciduous primocane leaves that are often palmately compound with five leaflets. Cultivated plants sometimes alternate from erect to creeping (or vice-versa) in successive years, depending on changing soil conditions. The following nothospecies names are based on putative hybrids involving Rubus setosus and: R. allegheniensis (R. ×abbrevians Blanchard, R. ×aculiferus Fernald, R. ×adenocaulis Fernald, R. ×angustifoliatus L. H. Bailey, R. ×ascendens Blanchard, R. ×atwoodii L. H. Bailey, R. ×clausenii L. H. Bailey, R. ×flavinanus Blanchard, R. ×frondisentis Blanchard, R. ×perspicuus L. H. Bailey, R. ×ravus L. H. Bailey); R. canadensis (R. ×miscix L. H. Bailey [based on R. ×peculiaris Blanchard (not R. peculiaris Sampaio)]); R. flagellaris (R. ×alter L. H. Bailey, R. ×arcuans Fernald & H. St. John, R. ×biformispinus Blanchard, R. ×bigelovianus L. H. Bailey, R. ×bracteolifer Fernald, R. ×jacens Blanchard var. specialis L. H. Bailey, R. ×laevior (L. H. Bailey) Fernald [based on R. permixtus var. laevior L. H. Bailey], R. ×multiformis Blanchard, R. ×setospinosus L. H. Bailey, R. ×severus Brainerd ex Fernald); R. hispidus (R. ×adjacens Fernald, R. ×grandidens L. H. Bailey, R. ×harmonicus L. H. Bailey, R. ×jacens Blanchard, R. ×parlinii L. H. Bailey, R. ×spiculosus Fernald, R. ×tardatus Blanchard, R. ×tholiformis Fernald, R. ×trifrons Blanchard, R. ×zaplutus L. H. Bailey); R. pensilvanicus (R. ×wisconsinensis L. H. Bailey). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Rubus saxatilis is defined by its creeping, unarmed or weakly armed stems, ternate leaves, and relatively small white petals. It has very weak prickles that detach easily, sometimes giving plants an unarmed appearance. The species is known in the flora area from only two sites in southern Greenland (T. W. Böcher 1938; D. B. Jensen and K. D. Christensen 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 53. | FNA vol. 9, p. 52. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus |
Sibling taxa | ||
Synonyms | R. apparatus, R. beatus, R. benneri, R. bicknellii, R. boottianus, R. condignus, R. deaneanus, R. discretus, R. dissensus, R. dissimilis, R. electus, R. exter, R. fulleri, R. groutianus, R. gulosus, R. hispidoides, R. hispidus var. suberectus, R. jejunus, R. junceus, R. junior, R. lawrencei, R. mediocris, R. navus, R. nocivus, R. notatus, R. ortivus, R. perinvisus, R. racemiger, R. regionalis, R. reravus, R. rotundior, R. schneideri, R. semisetosus, R. spectatus, R. stipulatus, R. uniformis, R. univocus, R. vermontanus, R. viridifrons, R. wheeleri | |
Name authority | Bigelow: Fl. Boston. ed. 2, 198. (1824) | Linnaeus: Sp. Pl. 1: 494. (1753) |
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