Rubus setosus |
Rosaceae tribe Rubeae |
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bristly blackberry, ronce sétuleuse, setose blackberry |
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Habit | Shrubs, 2–10(–15) dm, armed. | Shrubs, subshrubs, or herbs, perennial; armed or unarmed. |
Stems | biennial, erect to arching, rarely creeping, not node- or tip-rooting, sparsely to moderately hairy, sparsely to densely long-, rarely short-stipitate-glandular, glands rounded to flattened, not pruinose; prickles absent or sparsely to moderately dense, erect to retrorse, weak, slender, 1–3(–5) mm, narrow-based; bristles sparse to dense, erect to retrorse, green to reddish, narrow, semirigid, sometimes flexible, not gland-tipped. |
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Leaves | deciduous, ternate or palmately compound, sometimes ± lustrous; stipules filiform to lanceolate, 5–20(–38) mm; leaflets (3–)5, terminal obovate or elliptic to rhombic, 4.5–11.5 × 2–7 cm, base cuneate to narrowly rounded, unlobed, margins moderately, coarsely serrate to doubly serrate or serrate-dentate, apex acute or acuminate to short-attenuate, abaxial surfaces unarmed or with bristles on midvein, sparsely to moderately hairy, eglandular or sparsely to moderately sessile- to long-stipitate-glandular along larger veins. |
alternate, imparipinnately or palmately compound or simple; stipules persistent [deciduous], free or adnate to petiole; venation pinnate or palmate. |
Inflorescences | terminal on short shoots, sometimes appearing axillary, 5–15(–20)-flowered, racemiform, cymiform, or thyrsiform. |
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Pedicels | prickles or bristles sparse to dense, erect to retrorse, moderately to densely hairy, moderately to densely stipitate-glandular. |
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Flowers | bisexual; petals white, obovate to oblanceolate, 6–13 mm; filaments filiform; ovaries glabrous. |
perianth and androecium perigynous; epicalyx bractlets absent; hypanthium flat to hemispheric; torus flat or convex to conic; carpels 5–150, styles apical, distinct; ovules 2, apical, collateral, only 1 maturing. |
Fruits | black, globose, 0.7–1.5 cm; drupelets 5–25, strongly coherent, separating with torus attached. |
aggregated drupelets; styles persistent, not elongate. |
2n | = 14, 21, 28, 35. |
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Rubus setosus |
Rosaceae tribe Rubeae |
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Phenology | Flowering Jun–Jul(–Aug). | |
Habitat | Open woodlands, savannas, prairies, meadows, disturbed areas, dry to wet soil | |
Elevation | 0–1000 m (0–3300 ft) | |
Distribution |
CT; DC; DE; IA; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; NB; NF; NS; ON; PE; QC
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North America; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands; Australia [Introduced nearly worldwide] |
Discussion | Rubus setosus is identified by erect to arching, rarely creeping, not tip-rooting, terete primocanes, always setose but variable in density, and deciduous primocane leaves that are often palmately compound with five leaflets. Cultivated plants sometimes alternate from erect to creeping (or vice-versa) in successive years, depending on changing soil conditions. The following nothospecies names are based on putative hybrids involving Rubus setosus and: R. allegheniensis (R. ×abbrevians Blanchard, R. ×aculiferus Fernald, R. ×adenocaulis Fernald, R. ×angustifoliatus L. H. Bailey, R. ×ascendens Blanchard, R. ×atwoodii L. H. Bailey, R. ×clausenii L. H. Bailey, R. ×flavinanus Blanchard, R. ×frondisentis Blanchard, R. ×perspicuus L. H. Bailey, R. ×ravus L. H. Bailey); R. canadensis (R. ×miscix L. H. Bailey [based on R. ×peculiaris Blanchard (not R. peculiaris Sampaio)]); R. flagellaris (R. ×alter L. H. Bailey, R. ×arcuans Fernald & H. St. John, R. ×biformispinus Blanchard, R. ×bigelovianus L. H. Bailey, R. ×bracteolifer Fernald, R. ×jacens Blanchard var. specialis L. H. Bailey, R. ×laevior (L. H. Bailey) Fernald [based on R. permixtus var. laevior L. H. Bailey], R. ×multiformis Blanchard, R. ×setospinosus L. H. Bailey, R. ×severus Brainerd ex Fernald); R. hispidus (R. ×adjacens Fernald, R. ×grandidens L. H. Bailey, R. ×harmonicus L. H. Bailey, R. ×jacens Blanchard, R. ×parlinii L. H. Bailey, R. ×spiculosus Fernald, R. ×tardatus Blanchard, R. ×tholiformis Fernald, R. ×trifrons Blanchard, R. ×zaplutus L. H. Bailey); R. pensilvanicus (R. ×wisconsinensis L. H. Bailey). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genus 1, species 250–700 (37 in the flora). The base chromosome number for Rubeae is x = 7. The tribe is host to Phragmidium rusts. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 53. | FNA vol. 9, p. 27. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | R. apparatus, R. beatus, R. benneri, R. bicknellii, R. boottianus, R. condignus, R. deaneanus, R. discretus, R. dissensus, R. dissimilis, R. electus, R. exter, R. fulleri, R. groutianus, R. gulosus, R. hispidoides, R. hispidus var. suberectus, R. jejunus, R. junceus, R. junior, R. lawrencei, R. mediocris, R. navus, R. nocivus, R. notatus, R. ortivus, R. perinvisus, R. racemiger, R. regionalis, R. reravus, R. rotundior, R. schneideri, R. semisetosus, R. spectatus, R. stipulatus, R. uniformis, R. univocus, R. vermontanus, R. viridifrons, R. wheeleri | |
Name authority | Bigelow: Fl. Boston. ed. 2, 198. (1824) | Dumortier: Anal. Fam. Pl., 39. (1829) |
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