Rubus pubescens
Rubus pensilvanicus
dwarf raspberry, dwarf red blackberry, dwarf red blackberry or raspberry or red raspberry, dwarf red blackberry raspberry, dwarf red raspberry, five-leaf bramble, red raspberry, ronce pubescente, trailing raspberry
Pennsylvania blackberry, ronce de pennsylvanie
creeping, flowering branches erect, glabrous or sparsely to moderately hairy, eglandular, not pruinose.
biennial, erect to arching, glabrous or sparsely to densely hairy, eglandular or sparsely to moderately, rarely densely, sessile- to short-stipitate-glandular, not pruinose;
prickles sparse to dense, erect or slightly retrorse, stout, 4–10 mm, broad-based.
deciduous, usually ternate, sometimes pedately 5-foliate;
stipules lance-ovate to oblanceolate, (5–)8–12(–20) mm;
leaflets 3(–5), petiolule of terminal leaflet 2–7 mm, terminal rhombic to obovate, (2–)4–8(–10) × 2–4(–6) cm, base cuneate, unlobed or 2-lobed, margins deeply serrate to doubly serrate, apex acute or acuminate, abaxial surfaces glabrous or sparsely hairy, eglandular.
deciduous, palmately compound, not lustrous;
stipules filiform to narrowly lanceolate, (3–)5–15(–20) mm;
leaflets (3–)5(–7), terminal ovate to lanceolate, 5–15 × 3–13 cm, base rounded to shallowly cordate, unlobed, margins finely to coarsely singly or doubly serrate, apex acuminate to long-attenuate, abaxial surfaces green, usually with retrorse prickles on midveins, moderately hairy, eglandular or sparsely to moderately sessile- to short-stipitate-glandular along veins.
1–3-flowered, sometimes umbelliform.
terminal on short shoots, usually appearing axillary, (2–)5–12(–16)-flowered, cymiform, racemiform, or thyrsiform.
moderately to densely long-hairy, eglandular to densely stipitate-glandular.
unarmed or prickles sparse, erect, glabrous or sparsely to densely hairy, eglandular or sparsely to moderately sessile- to short-stipitate-glandular.
bisexual;
petals white to pink, oblanceolate to obovate, (4–)6–8(–10) mm;
filaments laminar;
ovaries glabrous, styles glabrous.
bisexual;
petals white, usually obovate to elliptic, rarely suborbiculate, 8–40 mm;
filaments filiform;
ovaries glabrous.
red, globose to conical, 0.5–1.4 cm diam., drupelets (5–)10–25, loosely coherent, falling separately or as a unit with torus attached.
black, globose to cylindric, 1–2 cm;
drupelets 10–100, strongly coherent, separating with torus attached.
= 14.
= 14, 21, 28, 35, 36, 42.
Rubus pubescens
Rubus pensilvanicus
Rubus pubescens is recognized by its creeping, unarmed stems, oblanceolate to lance-ovate stipules, relatively small flowers, and white to pink petals. It hybridizes with R. arcticus subsp. acaulis in areas of sympatry, and the hybrids are referable to R. ×paracaulis L. H. Bailey [= R. pubescens var. paracaulis (L. H. Bailey) B. Boivin]. The hybrids are similar to R. arcticus subsp. acaulis in having more obovate, rounded leaflets, and larger, pink to magenta petals; they are larger, creeping, and have hairy and stipitate-glandular pedicels and sepals like R. pubescens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
As circumscribed here, Rubus pensilvanicus represents sect. Arguti (L. H. Bailey) L. H. Bailey, in which Bailey (1941–1945) recognized 109 species.
More strongly glandular plants of Rubus pensilvanicus may represent introgression with R. allegheniensis or other species, especially when exhibiting long-stipitate glands. Thyrsiform inflorescences are present in plants that have been called R. missouricus. There are reports (T. S. Mallah 1954; L. V. Clark and M. Jasieniuk 2012; B. Sutherland and L. A. Alice, unpubl.) of R. pensilvanicus hybridizing with R. ursinus.
Rubus pensilvanicus is introduced in California, Idaho, and Washington.
The following nothospecies names are based on putative hybrids involving Rubus pensilvanicus and: R. allegheniensis (R. ×avipes L. H. Bailey, R. ×floricomus Blanchard, R. ×orarius Blanchard); R. canadensis (R. ×amabilis Blanchard [not Focke], R. ×amicalis Blanchard, R. ×elegantulus Blanchard [=R. canadensis var. elegantulus (Blanchard) Farwell], R. ×multilicius L. H. Bailey, R. ×noveboracus L. H. Bailey, R. ×pergratus Blanchard [= R. canadensis var. pergratus (Blanchard) L. H. Bailey], R. ×crux Ashe); R. cuneifolius (R. ×acer L. H. Bailey, R. ×acer var. subacer L. H. Bailey, R. ×argutinus L. H. Bailey, R. ×floridensis L. H. Bailey); R. flagellaris (R. ×akermanii Fernald, R. ×darlingtonii L. H. Bailey, R. ×dissitiflorus Fernald, R. ×hypolasius Fernald, R. ×janssonii L. H. Bailey, R. ×largus L. H. Bailey, R. ×recurvicaulis Blanchard, R. rossbergianus Blanchard); R. setosus (R. ×wisconsinensis L. H. Bailey).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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