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flowering raspberry, purple flowering raspberry, ronce odorante

Chesapeake blackberry, Nanticoke blackberry, topsy

Habit Shrubs, 10–20 dm, unarmed. Shrubs, 7–30 dm, armed.
Stems

erect, sparsely to moderately hairy, moderately to densely stipitate-glandular, glands dark purple, not pruinose.

biennial, arching, sparsely to densely hairy, sparsely to densely sessile-glandular, rarely short-stipitate-glandular, not pruinose;

bark not papery, peeling;

prickles moderate to dense, hooked, retrorse, or erect, stout, 4–10 mm, broad- or narrow-based.

Leaves

deciduous, simple;

stipules lanceolate to ovate, 5–15 mm;

blade subrotund to reniform, 9–20(–30) × (10–)15–25(–30) cm, base cordate, palmately, ± deeply, (3–)5-lobed, margins finely, irregularly serrate to doubly serrate, apex acute to acuminate, abaxial surfaces sparsely to moderately hairy, sparsely to densely stipitate-glandular, glands dark purple.

deciduous to semievergreen, palmately compound, rarely ternate;

stipules filiform to linear, rarely narrowly lanceolate, 5–16(–20) mm;

leaflets 3–5, terminal broadly elliptic or ovate, (3.5–)4.5–8.5 × 2.5–6 cm, base rounded, unlobed, margins moderately to coarsely doubly, rarely singly, serrate, apex acute or short-acuminate, abaxial surfaces with hooked prickles on larger veins, short-velutinous to tomentose, sparsely to densely sessile-glandular, rarely short-stipitate-glandular.

Inflorescences

terminal and axillary, 4–7(–22)-flowered, cymiform to thyrsiform.

terminal and axillary, 3–15(–25)-flowered, cymiform to thyrsiform, not projected well beyond subtending leaves.

Pedicels

pubescent, moderately to densely stipitate-glandular, glands dark purple.

prickles moderate to dense, hooked, retrorse, or erect, densely hairy, eglandular or sparsely to moderately sessile-glandular.

Flowers

bisexual;

petals usually magenta, rarely white, broadly obovate to suborbiculate, (12–)17–25(–30) mm;

filaments filiform;

ovaries distally densely hairy, styles clavate, glabrous.

bisexual;

petals white to pale pink, obovate or elliptic to orbiculate, (8–)10–15 mm;

filaments filiform;

ovaries glabrous or apically hairy.

Fruits

pale to dark red, hemispheric, 0.7–1.5 cm;

drupelets 30–60, coherent, separating from torus.

black, globose to subcylindric, 1–2 cm;

drupelets (15–)20–40(–50), strongly coherent, separating with torus attached.

2n

= 14.

= 21, 28.

Rubus odoratus

Rubus pascuus

Phenology Flowering May–Jul. Flowering (Apr–)May–Jun.
Habitat Moist shady sites in deciduous forests, margins of woods, rocky slopes, wooded talus, stream banks, roadsides Woodland edges, swamp margins, dry thickets, open and often disturbed areas, roadsides
Elevation 10–1500 m (0–4900 ft) 0–300(–700) m (0–1000(–2300) ft)
Distribution
from FNA
AL; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WA; WI; WV; NB; ON; QC [Introduced in Europe]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DE; GA; IL; IN; KY; MD; MO; MS; NC; NJ; OK; PA; SC; TN; TX; VA
[BONAP county map]
Discussion

Rubus odoratus is introduced in Washington State. The species is distinguished from other flowering raspberries by its erect, unarmed stems, simple leaves, large flowers, magenta petals, glabrous, clavate styles, and purple stipitate glands densely covering most plant parts. A hybrid with R. nutkanus (R. ×fraseri Rehder) is thought to occur in areas of overlap in northern Michigan (E. G. Voss 1972–1996, vol. 2).

The Cherokee used leaf infusions for labor pains and the Iroquois used plant infusions for miscarriage (P. Bergner 1997). Rubus odoratus is grown as an ornamental for its relatively large flowers and magenta petals and its edible fruits that are somewhat dry and insipid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus pascuus is probably more widespread than indicated here. The species is most likely to be mistaken for R. bifrons, R. cuneifolius, or R. pensilvanicus. Unlike R. bifrons, R. pascuus is generally more densely glandular, has much shorter, compact inflorescences that usually barely surpass the leaves, and tends to hold its primocane leaves relatively erect (versus spreading in R. bifrons). Although both can be semievergreen, R. pascuus will lose its leaves more readily than R. bifrons in areas where they co-occur. When compared with R. cuneifolius, R. pascuus has pale pink to white petals and ovate to broadly elliptic leaflets with flat margins, the entire part restricted to the base; R. cuneifolius has white petals and cuneate to obovate leaflets with usually revolute margins, the proximal third entire. When compared with R. pensilvanicus, R. pascuus has pale pink to white petals. Rubus pensilvanicus only has white petals as well as abaxial leaf surfaces, which are white to gray-green in R. pascuus with dense, short-velutinous to tomentose indument, while those of R. pensilvanicus can be hairy but never whitish to gray-green. A notable aspect of R. pascuus is how vigorously armed it is, more so than most other Rubus species in the flora area.

Rubus pascuus may represent a stabilized hybrid between a European and a native blackberry species, suggested by whitish pubescence on the abaxial leaf surface and pale pink petals. L. H. Bailey placed R. pascuus in his sect. Cuneifolii (L. H. Bailey) L. H. Bailey, likely because of the whitish hairs on the abaxial leaf surface.

The use of the name Rubus pascuus here is tentative; it is the best fit to the material observed. This treatment follows H. A. Davis (1990) for the choice of the name to use and its associated synonymy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 47. FNA vol. 9, p. 49.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Synonyms Rubacer odoratum, R. odoratus var. albidus, R. odoratus var. columbianus
Name authority Linnaeus: Sp. Pl. 1: 494. (1753) L. H. Bailey: Gentes Herb. 5: 440, fig. 200. (1943)
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