Rubus nutkanus |
Rosaceae tribe Rubeae |
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thimbleberry |
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Habit | Shrubs, 5–20(–30) dm, unarmed. | Shrubs, subshrubs, or herbs, perennial; armed or unarmed. |
Stems | biennial, erect, sparsely hairy, glabrescent, moderately stipitate-glandular, glands yellowish to reddish, not pruinose. |
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Leaves | deciduous, simple; stipules lanceolate to ovate, 5–15 mm; blade orbiculate to reniform, 5–20 × 5–25 cm, base cordate, palmately, shallowly to moderately deeply, (3–)5(–7)-lobed, margins coarsely, irregularly serrate to doubly serrate, apex shortly acuminate to obtuse, abaxial surfaces glabrate to densely hairy, sparsely to moderately stipitate-glandular, glands yellowish to reddish. |
alternate, imparipinnately or palmately compound or simple; stipules persistent [deciduous], free or adnate to petiole; venation pinnate or palmate. |
Inflorescences | terminal and axillary, (1–)3–7(–15), cymiform to thyrsiform. |
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Pedicels | sparsely to moderately hairy, moderately to densely stipitate-glandular, glands yellowish to reddish. |
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Flowers | bisexual; petals white, broadly obovate, (10–)14–22(–28) mm; filaments filiform; ovaries distally densely hairy, styles clavate, glabrous. |
perianth and androecium perigynous; epicalyx bractlets absent; hypanthium flat to hemispheric; torus flat or convex to conic; carpels 5–150, styles apical, distinct; ovules 2, apical, collateral, only 1 maturing. |
Fruits | red, hemispheric, 1–1.8 cm; drupelets 50–60, coherent, separating from torus. |
aggregated drupelets; styles persistent, not elongate. |
2n | = 14. |
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Rubus nutkanus |
Rosaceae tribe Rubeae |
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Phenology | Flowering May–Aug. | |
Habitat | Moist areas, open woods, thickets, clearings, stream banks, canyons, grassy meadows, rocky cliffs, sand dunes, upper beaches, dry sandy areas, roadsides | |
Elevation | 0–3000 m (0–9800 ft) | |
Distribution |
AK; AZ; CA; CO; IA; ID; IL; MI; MN; MT; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; ON; Mexico (Chihuahua) |
North America; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands; Australia [Introduced nearly worldwide] |
Discussion | Rubus nutkanus is distinguished from other flowering raspberries by its erect, unarmed stems, simple leaves, large flowers, white petals, glabrous, clavate styles, and yellowish to reddish stipitate glands covering most plant parts. A hybrid with R. odoratus (R. ×fraseri Rehder) is thought to occur in areas of overlap in northern Michigan (E. G. Voss 1972–1996, vol. 2). Varieties and forms have been described based on indument and glandularity of stems, leaves, petioles, pedicels, and sepals (for example, M. L. Fernald 1950). Different variants often occur mixed in the same population and generally do not correspond to definite geographic regions (N. C. Fassett 1941). Plants from coastal California (considered as var. velutinus) tend to have densely hairy leaf abaxial surfaces; this variation may be environmentally induced; other Rubus species show the same pattern (for example, R. spectabilis var. franciscanus) in that area. Wild thimbleberries are collected in Keweenaw Peninsula, Michigan, and are used in beverages and especially for jam. Rubus nutkanus is typically associated with western North America but was originally described from plants at its eastern range limit in northern Michigan. The Great Lakes populations are disjunct from western mountain populations by around 600 miles. The name Rubus parviflorus var. grandiflorus Farwell, which applies here, is illegitimate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genus 1, species 250–700 (37 in the flora). The base chromosome number for Rubeae is x = 7. The tribe is host to Phragmidium rusts. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 48. | FNA vol. 9, p. 27. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | R. parviflorus | |
Name authority | Mociño ex Seringe: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 2: 566. (1825) | Dumortier: Anal. Fam. Pl., 39. (1829) |
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