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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry

European blackberry

Habit Shrubs, to 1.5 dm, armed. Shrubs, 10–20 dm, armed.
Stems

perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose;

prickles sparse, strongly retrorse, stout, to 1 mm, broad-based.

biennial, arching, usually densely, sometimes moderately, hairy, usually eglandular, sometimes sessile- to short-stipitate-glandular, not pruinose;

prickles moderate to dense, recurved to erect, slender, 3–10 mm, broad-based.

Leaves

evergreen, simple or ternate;

stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm;

blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh.

deciduous or ± persistent, palmately compound;

stipules filiform to linear, 5–15 mm;

leaflets 3–5, terminal usually suborbiculate to orbiculate, sometimes broadly elliptic, 4–9 × 4–8 cm, base rounded to shallowly cordate, unlobed, margins moderately to coarsely serrate, apex acute to cuspidate, abaxial surfaces with hooked prickles on midvein, densely white-gray to gray-green tomentose, eglandular or obscurely short-stipitate- to sessile-glandular along midveins.

Inflorescences

1–2-flowered.

terminal, 7–35-flowered, cymiform or thyrsiform.

Pedicels

prickles scattered, retrorse, moderately to densely hairy, eglandular.

prickles moderate to dense, recurved to erect, densely hairy, moderately to densely stipitate-glandular.

Flowers

bisexual;

petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm;

filaments filiform;

ovaries moderately hairy, styles glabrous.

bisexual;

petals pink to magenta, elliptic to suborbiculate, 6–12(–15) mm;

filaments filiform;

ovaries apically hairy.

Fruits

red, hemispheric, 0.4–1 cm;

drupelets 3–10, not coherent, separating from torus.

black, globose, 1–1.5 cm;

drupelets 15–40, coherent, separating with torus attached.

2n

= 14.

= 28.

Rubus nivalis

Rubus vestitus

Phenology Flowering Jun–Sep. Flowering Jul–Sep.
Habitat Moist, semishaded forests, glades, moist soil, logged areas Woodland edges, open areas, disturbed areas, in moist soil
Elevation 0–1700 m (0–5600 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; BC; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008).

The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936).

The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus vestitus can be distinguished, especially from the closely related R. bifrons and R. ulmifolius, by its long-stipitate-glandular hairs, particularly in the inflorescence, and terminal primocane leaflets that are typically suborbiculate and abaxially densely tomentose. Rubus allegheniensis can possess similar glandular hairs; it does not have pink petals, thyrsiform inflorescences, and rounded and abaxially densely tomentose, often whitened terminal leaflets.

Rubus vestitus has not yet been found in Washington, but it is likely there. Because of the overall similarity of R. vestitus to R. bifrons, it is possible that this potentially weedy species is widespread, but rare and possibly overlooked, occurring as far south as California. In Oregon, R. vestitus prefers shady habitats (B. Wilson, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 45. FNA vol. 9, p. 56.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus
Name authority Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) Weihe & Nees: in M. J. Bluff et al., Comp. Fl. German. 1: 684. (1825)
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