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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry

Japanese wineberry, wine raspberry, wineberry

Habit Shrubs, to 1.5 dm, armed. Shrubs, 10–20(–30) dm, armed.
Stems

perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose;

prickles sparse, strongly retrorse, stout, to 1 mm, broad-based.

biennial, arching, sparsely to moderately hairy, long-stipitate-glandular, not pruinose;

prickles sparse to dense, erect to hooked, slender, 2–8(–10) mm, slender-based;

bristles dense, 2–6 mm, usually gland-tipped and reddish purple, glands ellipsoid to narrowly cup-shaped.

Leaves

evergreen, simple or ternate;

stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm;

blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh.

deciduous, ternate;

stipules filiform, 5–12 mm;

terminal leaflets broadly deltate or ovate to suborbiculate, 4–15 × 3.5–14 cm, base truncate, rounded, or shallowly cordate, unlobed or dentate to shallowly, sharply or bluntly 3-lobed, margins coarsely serrate or doubly serrate, apex acute to acuminate, abaxial surfaces with scattered, erect to curved prickles on moderate- to large-sized veins, densely white-hairy, moderately to densely long-stipitate-glandular along veins.

Inflorescences

1–2-flowered.

terminal and axillary, 5–30-flowered, cymiform to racemiform.

Pedicels

prickles scattered, retrorse, moderately to densely hairy, eglandular.

usually unarmed or prickles sparse, erect, moderately to densely hairy, densely long-stipitate-glandular.

Flowers

bisexual;

petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm;

filaments filiform;

ovaries moderately hairy, styles glabrous.

bisexual;

petals white to pink, ovate to suborbiculate, 4–6 mm;

filaments laminar;

ovaries glabrous.

Fruits

red, hemispheric, 0.4–1 cm;

drupelets 3–10, not coherent, separating from torus.

red to maroon, globose, 1–1.5 cm;

drupelets 15–40, strongly coherent, separating from torus.

2n

= 14.

= 14.

Rubus nivalis

Rubus phoenicolasius

Phenology Flowering Jun–Sep. Flowering May–Jul.
Habitat Moist, semishaded forests, glades, moist soil, logged areas Woodlands, roadsides, disturbed open areas, moist soil
Elevation 0–1700 m (0–5600 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IL; IN; KY; MA; MD; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV; BC; e Asia (China, Japan, Korea) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008).

The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936).

The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus phoenicolasius was introduced to North America for edible fruit, breeding stock, and for ornament. Attractive for its typically reddish purple glandular hairs, R. phoenicolasius nonetheless can be invasive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 45. FNA vol. 9, p. 51.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Name authority Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) Maximowicz: Bull. Acad. Imp. Sci. Saint-Pétersbourg 17: 160. (1872)
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