Rubus nivalis |
Rubus phoenicolasius |
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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry |
Japanese wineberry, wine raspberry, wineberry |
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Habit | Shrubs, to 1.5 dm, armed. | Shrubs, 10–20(–30) dm, armed. |
Stems | perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose; prickles sparse, strongly retrorse, stout, to 1 mm, broad-based. |
biennial, arching, sparsely to moderately hairy, long-stipitate-glandular, not pruinose; prickles sparse to dense, erect to hooked, slender, 2–8(–10) mm, slender-based; bristles dense, 2–6 mm, usually gland-tipped and reddish purple, glands ellipsoid to narrowly cup-shaped. |
Leaves | evergreen, simple or ternate; stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm; blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh. |
deciduous, ternate; stipules filiform, 5–12 mm; terminal leaflets broadly deltate or ovate to suborbiculate, 4–15 × 3.5–14 cm, base truncate, rounded, or shallowly cordate, unlobed or dentate to shallowly, sharply or bluntly 3-lobed, margins coarsely serrate or doubly serrate, apex acute to acuminate, abaxial surfaces with scattered, erect to curved prickles on moderate- to large-sized veins, densely white-hairy, moderately to densely long-stipitate-glandular along veins. |
Inflorescences | 1–2-flowered. |
terminal and axillary, 5–30-flowered, cymiform to racemiform. |
Pedicels | prickles scattered, retrorse, moderately to densely hairy, eglandular. |
usually unarmed or prickles sparse, erect, moderately to densely hairy, densely long-stipitate-glandular. |
Flowers | bisexual; petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm; filaments filiform; ovaries moderately hairy, styles glabrous. |
bisexual; petals white to pink, ovate to suborbiculate, 4–6 mm; filaments laminar; ovaries glabrous. |
Fruits | red, hemispheric, 0.4–1 cm; drupelets 3–10, not coherent, separating from torus. |
red to maroon, globose, 1–1.5 cm; drupelets 15–40, strongly coherent, separating from torus. |
2n | = 14. |
= 14. |
Rubus nivalis |
Rubus phoenicolasius |
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Phenology | Flowering Jun–Sep. | Flowering May–Jul. |
Habitat | Moist, semishaded forests, glades, moist soil, logged areas | Woodlands, roadsides, disturbed open areas, moist soil |
Elevation | 0–1700 m (0–5600 ft) | 0–1200 m (0–3900 ft) |
Distribution |
CA; ID; OR; WA; BC
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AL; AR; CT; DC; DE; GA; IL; IN; KY; MA; MD; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV; BC; e Asia (China, Japan, Korea) [Introduced in North America]
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Discussion | Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008). The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936). The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Rubus phoenicolasius was introduced to North America for edible fruit, breeding stock, and for ornament. Attractive for its typically reddish purple glandular hairs, R. phoenicolasius nonetheless can be invasive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 45. | FNA vol. 9, p. 51. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus |
Sibling taxa | ||
Name authority | Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) | Maximowicz: Bull. Acad. Imp. Sci. Saint-Pétersbourg 17: 160. (1872) |
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