Rubus nivalis |
Rubus pedatus |
|
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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry |
creeping raspberry, five leaf dwarf bramble, five-leaf bramble, strawberry bramble, strawberry dwarf bramble, strawberry-leaf raspberry |
|
Habit | Shrubs, to 1.5 dm, armed. | Herbs, to 1 dm, unarmed. |
Stems | perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose; prickles sparse, strongly retrorse, stout, to 1 mm, broad-based. |
creeping, flowering branches erect, sparsely hairy, eglandular or sparsely stipitate-glandular, not pruinose. |
Leaves | evergreen, simple or ternate; stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm; blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh. |
deciduous, pedately compound; stipules broadly ovate to suborbiculate, 2–3 mm; leaflets 3 (often appearing as if 5), lateral leaflets deeply lobed, sinuses nearly to leaflet base, terminal obovate to obovate-rhombic, (1–)1.5–2.5(–3.4) × (0.8–)1.2–2(–2.5) cm, base cuneate, margins coarsely singly or doubly serrate, apex rounded to obtuse, abaxial surfaces glabrous or sparsely hairy on midvein, eglandular or sparsely stipitate-glandular. |
Inflorescences | 1–2-flowered. |
1-flowered. |
Pedicels | prickles scattered, retrorse, moderately to densely hairy, eglandular. |
sparsely hairy, eglandular or sparsely stipitate-glandular. |
Flowers | bisexual; petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm; filaments filiform; ovaries moderately hairy, styles glabrous. |
bisexual; petals white, oblong to narrowly obovate, 6–10 mm; filaments filiform; ovaries glabrous, styles glabrous. |
Fruits | red, hemispheric, 0.4–1 cm; drupelets 3–10, not coherent, separating from torus. |
red, 0.6–1 cm; drupelets 3–6, weakly coherent, separating from torus. |
2n | = 14. |
= 14. |
Rubus nivalis |
Rubus pedatus |
|
Phenology | Flowering Jun–Sep. | Flowering Jun–Aug. |
Habitat | Moist, semishaded forests, glades, moist soil, logged areas | Coniferous woods, forest edges, glades, meadows, bogs, stream banks, streambeds, roadsides |
Elevation | 0–1700 m (0–5600 ft) | 20–2300 m (100–7500 ft) |
Distribution |
CA; ID; OR; WA; BC
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AK; ID; MT; OR; WA; AB; BC; YT; ne Asia
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Discussion | Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008). The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936). The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Rubus pedatus is recognized by its creeping, unarmed stems, pedately 3–5-foliate leaves, relatively small flowers, white petals, and glabrous ovaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 45. | FNA vol. 9, p. 49. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus |
Sibling taxa | ||
Name authority | Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) | Smith: Pl. Icon. Ined. 3: plate 63. (1791) |
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