Rubus nivalis |
Rubus leucodermis |
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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry |
black raspberry, blackcap, blackcap raspberry, dark raspberry, dwarf bramble, western black raspberry, white-bark raspberry |
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Habit | Shrubs, to 1.5 dm, armed. | Shrubs, 10–30 dm, armed. | ||||
Stems | perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose; prickles sparse, strongly retrorse, stout, to 1 mm, broad-based. |
biennial, ascending, often arching, glabrous or sparsely puberulent, eglandular or sparsely to densely short-stipitate-glandular, strongly pruinose; prickles moderate to dense, erect or curved, slender to stout, 4–8 mm, usually broad-based, sometimes narrow-based. |
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Leaves | evergreen, simple or ternate; stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm; blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh. |
deciduous, palmately compound or ternate; stipules filiform, 5–10 mm; petiole with hooked or erect prickles, glabrous, eglandular or sparsely to densely sessile- to short-stipitate-glandular; lateral leaflets stalked, at least 2 larger ones, often with prickles, glabrous, eglandular or stipitate-glandular; leaflets 3–5(–7), terminal ovate to lanceolate, 6–10 × 3–6 cm, base cordate to truncate, unlobed or shallowly 3-lobed, margins serrate to doubly serrate, apex acuminate, abaxial surface sometimes with prickles on midrib and some lateral veins, glabrous to strongly white-tomentose, usually eglandular. |
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Inflorescences | 1–2-flowered. |
axillary and terminal, (1–)3–10(–12)-flowered, flat-topped cymiform or umbelliform. |
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Pedicels | prickles scattered, retrorse, moderately to densely hairy, eglandular. |
prickles sparse to dense, hooked or erect, puberulent, eglandular or sparsely to densely sessile- or short-stipitate-glandular. |
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Flowers | bisexual; petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm; filaments filiform; ovaries moderately hairy, styles glabrous. |
bisexual; petals erect, then ascending, white, oblong to oblanceolate, 3–6(–8) mm; filaments laminar; ovaries pubescent, rarely glabrous. |
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Fruits | red, hemispheric, 0.4–1 cm; drupelets 3–10, not coherent, separating from torus. |
reddish purple to nearly black, depressed globose to conic, 1–2 cm; drupelets 20–60, coherent, separating from torus. |
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2n | = 14. |
= 14. |
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Rubus nivalis |
Rubus leucodermis |
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Phenology | Flowering Jun–Sep. | |||||
Habitat | Moist, semishaded forests, glades, moist soil, logged areas | |||||
Elevation | 0–1700 m (0–5600 ft) | |||||
Distribution |
CA; ID; OR; WA; BC
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AK; AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; n Mexico
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Discussion | Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008). The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936). The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Rubus leucodermis is similar to R. occidentalis, differing primarily by its leaflets having acuminate apices and lateral leaflets that are distinctly stalked. Prickles on the pedicels of R. occidentalis are erect; those of R. leucodermis tend to be hooked. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 45. | FNA vol. 9, p. 44. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | R. occidentalis subsp. leucodermis | |||||
Name authority | Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) | Douglas ex Torrey & A. Gray: Fl. N. Amer. 1: 454. (1840) | ||||
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