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dwarf snow bramble, snow bramble, snow dewberry, snow dwarf bramble, snow raspberry

cut-leaf blackberry, cutleaf evergreen blackberry, evergreen blackberry

Habit Shrubs, to 1.5 dm, armed. Shrubs, 1–20 dm, armed.
Stems

perennial, creeping, sparsely hairy, glabrescent, eglandular, not pruinose;

prickles sparse, strongly retrorse, stout, to 1 mm, broad-based.

biennial, arching, sometimes creeping, glabrate to moderately hairy, sparsely to densely sessile- to short-stipitate-glandular, not or weakly pruinose;

prickles moderately dense to dense, suberect to retrorse or hooked, stout, 2–8 mm, broad-based.

Leaves

evergreen, simple or ternate;

stipules adnate to petioles, broadly elliptic to ovate, (6–)8–10 mm;

blade ovate to cordate, (2.5–)3–5(–8) cm, lobe or leaflet base cordate, shallowly 3-lobed, margins coarsely, singly or doubly dentate, apex acute to obtuse, abaxial surfaces with prickles along midveins, glabrous or sparsely hairy, eglandular, both surfaces lustrous fresh.

evergreen to late-deciduous, ternate or palmately compound;

stipules filiform, 5–15 mm;

leaflets 3–5, terminal ovate to elliptic, 6–12 × 7–12 cm, base rounded to cordate, usually deeply, rarely shallowly, lobed, margins coarsely serrate to doubly serrate, apex acuminate to short-attenuate, abaxial surfaces with hooked prickles primarily on midvein, moderately to densely hairy, eglandular or moderately sessile- to short-stipitate-glandular primarily on midvein.

Inflorescences

1–2-flowered.

terminal, 5–25-flowered, thyrsiform.

Pedicels

prickles scattered, retrorse, moderately to densely hairy, eglandular.

prickles moderate to dense, erect to reflexed, densely hairy, moderately to densely sessile- to short-stipitate-glandular.

Flowers

bisexual;

petals magenta to pink, elliptic to oblanceolate or spatulate, (5–)8–10 mm;

filaments filiform;

ovaries moderately hairy, styles glabrous.

bisexual;

petals white to pink, elliptic to obovate, 8–15 mm;

filaments filiform;

ovaries usually apically hairy.

Fruits

red, hemispheric, 0.4–1 cm;

drupelets 3–10, not coherent, separating from torus.

black, globose to subcylindric, 1–1.5 cm;

drupelets 25–60, strongly coherent, separating with torus attached.

2n

= 14.

= 28.

Rubus nivalis

Rubus laciniatus

Phenology Flowering Jun–Sep. Flowering Jun–Sep(–Oct).
Habitat Moist, semishaded forests, glades, moist soil, logged areas Woodland edges, semiopen woods, rocky and/or moist soil, open areas, often disturbed, roadsides
Elevation 0–1700 m (0–5600 ft) 0–1800 m (0–5900 ft)
Distribution
from FNA
CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; CT; DC; DE; ID; IL; IN; KY; MA; MD; MI; MO; MT; NC; NJ; NY; OH; OR; PA; RI; SC; TN; VA; VT; WA; WV; WY; BC; ON; Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Rubus nivalis is recognized by its creeping, prickly stems, simple to ternate, evergreen leaves, broadly elliptic to ovate stipules, two leaflets, relatively small flowers, and magenta to pink petals. Its closest relative is likely the Mexican R. pumilus Focke. Asian species previously classified in subg. Chamaebatus (Focke) Focke are hexaploid (M. M. Thompson 1997) and not phylogenetically close; R. nivalis appears to be sister to all blackberries of subg. Rubus (L. A. Alice and C. S. Campbell 1999; Alice et al. 2008).

The fruits of Rubus nivalis are eaten fresh, stewed, and canned by the Hoh and Quileute Indians (A. B. Reagan 1936).

The only known specimen of Rubus nivalis from California was collected in 1961 from Del Norte County at 1250 m near the Oregon border.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rubus laciniatus is the only blackberry in the flora area that has leaflets so deeply lobed and dissected or has 3-lobed petals. Rare individuals bearing relatively unlobed leaflets and perianth parts may be part of the natural variation of this species or be hybrids with some other species. Putative natural hybrids have been reported with R. bifrons (as R. procerus; R. K. Bammi and H. P. Olmo 1966). Such unusual plants of R. laciniatus would be distinguished from the native blackberries by their thyrsiform and notably stipitate-glandular inflorescences and evergreen to late-deciduous leaves.

Although widely introduced in the flora area, Rubus laciniatus is a less aggressive weed than R. bifrons.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 45. FNA vol. 9, p. 43.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Name authority Douglas: in W. J. Hooker, Fl. Bor.-Amer. 1: 181. (1832) Willdenow: Hort. Berol. 2: plate 82. (1806)
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