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Rubus bartonianus

Barton's raspberry, bartonberry

black raspberry, blackcap, blackcap raspberry, dark raspberry, dwarf bramble, western black raspberry, white-bark raspberry

Habit Shrubs, 8–25 dm, unarmed. Shrubs, 10–30 dm, armed.
Stems

erect, sparsely short-hairy, glabrescent, eglandular, not pruinose.

biennial, ascending, often arching, glabrous or sparsely puberulent, eglandular or sparsely to densely short-stipitate-glandular, strongly pruinose;

prickles moderate to dense, erect or curved, slender to stout, 4–8 mm, usually broad-based, sometimes narrow-based.

Leaves

deciduous, simple;

stipules lanceolate, 4–6 mm;

blade cordate to broadly ovate, (2–)2.5–4(–5) × (2.5–)3.5–4.5(–5.5) cm, base deeply cordate, 3–5-lobed, lobe apices acute to obtuse, margins coarsely doubly dentate, abaxial surfaces glabrous or sparsely hairy, eglandular or sparsely stipitate-glandular.

deciduous, palmately compound or ternate;

stipules filiform, 5–10 mm;

petiole with hooked or erect prickles, glabrous, eglandular or sparsely to densely sessile- to short-stipitate-glandular;

lateral leaflets stalked, at least 2 larger ones, often with prickles, glabrous, eglandular or stipitate-glandular;

leaflets 3–5(–7), terminal ovate to lanceolate, 6–10 × 3–6 cm, base cordate to truncate, unlobed or shallowly 3-lobed, margins serrate to doubly serrate, apex acuminate, abaxial surface sometimes with prickles on midrib and some lateral veins, glabrous to strongly white-tomentose, usually eglandular.

Inflorescences

1-flowered.

axillary and terminal, (1–)3–10(–12)-flowered, flat-topped cymiform or umbelliform.

Pedicels

moderately hairy, eglandular or sparsely stipitate-glandular.

prickles sparse to dense, hooked or erect, puberulent, eglandular or sparsely to densely sessile- or short-stipitate-glandular.

Flowers

bisexual;

petals white, obovate, (15–)20–25 mm;

filaments filiform;

ovaries glabrous, styles clavate, villous.

bisexual;

petals erect, then ascending, white, oblong to oblanceolate, 3–6(–8) mm;

filaments laminar;

ovaries pubescent, rarely glabrous.

Fruits

deep red, hemispheric, to 1 cm;

drupelets 10–30, coherent, separating from torus.

reddish purple to nearly black, depressed globose to conic, 1–2 cm;

drupelets 20–60, coherent, separating from torus.

2n

= 14.

Rubus bartonianus

Rubus leucodermis

Phenology Flowering Mar–May.
Habitat Dry, rocky slopes
Elevation 300–400 m [1000–1300 ft]
Distribution
from FNA
ID; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Rubus bartonianus is distinguished from the other flowering raspberries within its geographic range by its erect, unarmed stems, relatively small, simple leaves with acute to obtuse lobes, deeply cordate bases, sparsely hairy or glabrous abaxial surfaces, relatively large flowers with white petals, and densely long-hairy, clavate styles. The leaves superficially resemble those of Acer glabrum or some species of Ribes.

Rubus bartonianus is most similar to R. neomexicanus but especially R. deliciosus. The species is known only from the Snake River Canyon of Idaho and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Rubus leucodermis is similar to R. occidentalis, differing primarily by its leaflets having acuminate apices and lateral leaflets that are distinctly stalked. Prickles on the pedicels of R. occidentalis are erect; those of R. leucodermis tend to be hooked.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves abaxially white-tomentose; drupelets (20–)30–60.
R. leucodermis var. leucodermis
1. Leaves abaxially glabrous, rarely sparsely hairy; drupelets 20–40.
R. leucodermis var. nigerrimus
Source FNA vol. 9, p. 36. FNA vol. 9, p. 44.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus
Sibling taxa
R. allegheniensis, R. arcticus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. leucodermis, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
R. allegheniensis, R. arcticus, R. bartonianus, R. bifrons, R. caesius, R. canadensis, R. chamaemorus, R. cuneifolius, R. deliciosus, R. flagellaris, R. glaucifolius, R. hispidus, R. idaeus, R. illecebrosus, R. laciniatus, R. lasiococcus, R. neomexicanus, R. nivalis, R. niveus, R. nutkanus, R. occidentalis, R. odoratus, R. parviflorus, R. parvifolius, R. pascuus, R. pedatus, R. pensilvanicus, R. phoenicolasius, R. pubescens, R. repens, R. saxatilis, R. setosus, R. spectabilis, R. trivialis, R. ulmifolius, R. ursinus, R. vestitus
Subordinate taxa
R. leucodermis var. leucodermis, R. leucodermis var. nigerrimus
Synonyms R. occidentalis subsp. leucodermis
Name authority M. Peck: Rhodora 36: 267. (1934) Douglas ex Torrey & A. Gray: Fl. N. Amer. 1: 454. (1840)
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