Rubus allegheniensis |
Rubus pensilvanicus |
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alleghaney blackberry, Alleghany blackberry, Allegheny blackberry, common blackberry, prairie rose, ronce des alléghanys, sow-teat berry |
Pennsylvania blackberry, ronce de pennsylvanie |
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Habit | Shrubs, 10–30 dm, armed. | Shrubs, 10–30 dm, armed. |
Stems | biennial, erect-arching, sparsely to densely hairy, sparsely to abundantly stipitate-glandular, glands usually flattened to cupulate, not pruinose; prickles sparse to dense, erect or slightly retrorse, stout, 4–10 mm, broad-based. |
biennial, erect to arching, glabrous or sparsely to densely hairy, eglandular or sparsely to moderately, rarely densely, sessile- to short-stipitate-glandular, not pruinose; prickles sparse to dense, erect or slightly retrorse, stout, 4–10 mm, broad-based. |
Leaves | deciduous, palmately compound, not lustrous; stipules filiform to narrowly lanceolate, (2–)5–15(–20) mm; leaflets (3–)5(–7), terminal ovate to lanceolate, 7–16 × 4–9 cm, base rounded to shallowly cordate, unlobed, margins finely to coarsely serrate or doubly serrate, apex acuminate to long-attenuate, abaxial surfaces with hooked prickles on midveins, moderately hairy, moderately to densely stipitate-glandular on veins. |
deciduous, palmately compound, not lustrous; stipules filiform to narrowly lanceolate, (3–)5–15(–20) mm; leaflets (3–)5(–7), terminal ovate to lanceolate, 5–15 × 3–13 cm, base rounded to shallowly cordate, unlobed, margins finely to coarsely singly or doubly serrate, apex acuminate to long-attenuate, abaxial surfaces green, usually with retrorse prickles on midveins, moderately hairy, eglandular or sparsely to moderately sessile- to short-stipitate-glandular along veins. |
Inflorescences | terminal on short shoots, sometimes appearing axillary, (5–)15–25-flowered, racemiform, often elongate. |
terminal on short shoots, usually appearing axillary, (2–)5–12(–16)-flowered, cymiform, racemiform, or thyrsiform. |
Pedicels | unarmed or prickles sparse, hooked, densely hairy, densely short- to long-stipitate-glandular. |
unarmed or prickles sparse, erect, glabrous or sparsely to densely hairy, eglandular or sparsely to moderately sessile- to short-stipitate-glandular. |
Flowers | bisexual; petals white, obovate to elliptic, rarely suborbiculate, 8–20 mm; filaments filiform; ovaries glabrous or glabrate. |
bisexual; petals white, usually obovate to elliptic, rarely suborbiculate, 8–40 mm; filaments filiform; ovaries glabrous. |
Fruits | black, globose to cylindric, 1–2 cm; drupelets 20–100, strongly coherent, separating with torus attached. |
black, globose to cylindric, 1–2 cm; drupelets 10–100, strongly coherent, separating with torus attached. |
2n | = 14, 21, 28. |
= 14, 21, 28, 35, 36, 42. |
Rubus allegheniensis |
Rubus pensilvanicus |
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Phenology | Flowering May–Jul. | Flowering May–Jul. |
Habitat | Woodlands, savannas, prairies, meadows, rock outcrops, disturbed areas, dry to damp soil | Woodlands, savannas, prairies, fields, meadows, swamps, rock outcrops, sand dunes, sandy soil, disturbed areas, dry to wet soil |
Elevation | 0–1600 m (0–5200 ft) | 0–1400 m (0–4600 ft) |
Distribution |
AL; AR; CA; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; BC; NB; NS; ON; PE; QC
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AL; AR; CA; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WA; WI; WV; NB; NF; NS; ON; PE; QC [Introduced in Pacific Islands (Hawaii)]
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Discussion | Rubus allegheniensis consists of erect-arching, usually well-armed plants, usually densely stipitate-glandular with flattened to cupulate glands, particularly about the inflorescence. The Menominee, Meskwaki, and Potawatomi used root extracts of Rubus allegheniensis to treat eye irritation (H. H. Smith 1923, 1928, 1933). The Ojibwa used boiled extracts of the canes as a diuretic and a root extract for treating diarrhea (Smith 1932). Rubus allegheniensis is introduced in British Columbia and California, where rare and in at least California probably no more than a waif. Rubus montanus (Porter) Porter, a name that has been used for this species, is a later homonym of R. montanus Libert ex Lejeune. The following nothospecies names are based on putative hybrids involving Rubus allegheniensis and: R. canadensis (R. ×forestalis L. H. Bailey, R. ×immanis Ashe); R. flagellaris (R. ×boyntonii Ashe, R. ×fraternalis L. H. Bailey [based on R. ×fraternus Brainerd & Peitersen (not R. fraternus Gremli), R. ×licens L. H. Bailey, R. ×ostryifolius Rydberg); R. pensilvanicus (R. ×avipes L. H. Bailey, R. ×floricomus Blanchard, R. ×orarius Blanchard); R. setosus (R. ×abbrevians Blanchard, R. ×aculiferus Fernald, R. ×adenocaulis Fernald, R. ×angustifoliatus L. H. Bailey, R. ×ascendens Blanchard, R. ×atwoodii L. H. Bailey, R. ×clausenii L. H. Bailey, R. ×flavinanus Blanchard, R. ×frondisentis Blanchard, R. ×perspicuus L. H. Bailey, R. ×ravus L. H. Bailey). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As circumscribed here, Rubus pensilvanicus represents sect. Arguti (L. H. Bailey) L. H. Bailey, in which Bailey (1941–1945) recognized 109 species. More strongly glandular plants of Rubus pensilvanicus may represent introgression with R. allegheniensis or other species, especially when exhibiting long-stipitate glands. Thyrsiform inflorescences are present in plants that have been called R. missouricus. There are reports (T. S. Mallah 1954; L. V. Clark and M. Jasieniuk 2012; B. Sutherland and L. A. Alice, unpubl.) of R. pensilvanicus hybridizing with R. ursinus. Rubus pensilvanicus is introduced in California, Idaho, and Washington. The following nothospecies names are based on putative hybrids involving Rubus pensilvanicus and: R. allegheniensis (R. ×avipes L. H. Bailey, R. ×floricomus Blanchard, R. ×orarius Blanchard); R. canadensis (R. ×amabilis Blanchard [not Focke], R. ×amicalis Blanchard, R. ×elegantulus Blanchard [=R. canadensis var. elegantulus (Blanchard) Farwell], R. ×multilicius L. H. Bailey, R. ×noveboracus L. H. Bailey, R. ×pergratus Blanchard [= R. canadensis var. pergratus (Blanchard) L. H. Bailey], R. ×crux Ashe); R. cuneifolius (R. ×acer L. H. Bailey, R. ×acer var. subacer L. H. Bailey, R. ×argutinus L. H. Bailey, R. ×floridensis L. H. Bailey); R. flagellaris (R. ×akermanii Fernald, R. ×darlingtonii L. H. Bailey, R. ×dissitiflorus Fernald, R. ×hypolasius Fernald, R. ×janssonii L. H. Bailey, R. ×largus L. H. Bailey, R. ×recurvicaulis Blanchard, R. rossbergianus Blanchard); R. setosus (R. ×wisconsinensis L. H. Bailey). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 34. | FNA vol. 9, p. 50. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus | Rosaceae > subfam. Rosoideae > tribe Rubeae > Rubus |
Sibling taxa | ||
Synonyms | R. villosus var. montanus, R. acadiensis, R. allegheniensis var. calycosus, R. allegheniensis var. gravesii, R. allegheniensis var. neoscoticus, R. allegheniensis var. plausus, R. allegheniensis var. populifolius, R. alumnus, R. apianus, R. attractus, R. auroralis, R. campester, R. concameratus, R. fryei, R. glandicaulis, R. impos, R. inclinis, R. irregularis, R. licitus, R. montpelierensis, R. nigrobaccus, R. nuperus, R. paulus, R. pugnax, R. rosa, R. saltuensis, R. sceleratus | R. abactus, R. ablatus, R. abundiflorus, R. amnicola, R. andrewsianus, R. argutus, R. arvensis, R. associus, R. barbarus, R. bellobatus, R. betulifolius, R. blakei, R. bractealis, R. brainerdii, R. burnhamii, R. bushii, R. cardianus, R. condensiflorus, R. congruus, R. cupressorum, R. defectionis, R. densipubens, R. densissimus, R. facetus, R. fatuus, R. floridus, R. frondosus, R. gnarus, R. heterophyllus, R. humilior, R. impar, R. independens, R. insons, R. insulanus, R. interioris, R. jennisonii, R. jugosus, R. kelloggii, R. latifoliolus, R. laudatus, R. leggii, R. libratus, R. louisianus, R. missouricus, R. mollior, R. multispinus, R. oklahomus, R. originalis, R. paludivagus, R. pauper, R. pauxillus, R. penetrans, R. perfoliosus, R. perpauper, R. persistens, R. philadelphicus, R. praepes, R. pratensis, R. prestonensis, R. pubifolius, R. recurvans, R. saepescandens, R. sativus, R. subsolanus, R. subtractus, R. suus, R. tygartensis, R. ucetanus, R. virilis, R. vixargutus, R. wahlii, R. wiegandii |
Name authority | Porter: Bull. Torrey Bot. Club 23: 153. (1896) | Poiret: in J. Lamarck et al., Encycl. 6: 246. (1804) |
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