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bryum moss, capillary bryum, capillary thread-moss

Habit Plants small, bright green. Plants small to usually medium-sized, green or red-green.
Stems

0.5–1.5 cm, distinctly singly rosulate, innovations short, rosulate.

1.5–2 cm, distinctly singly rosulate, innovations short, rosulate.

Leaves

of main rosette and innovations similar, regularly spirally twisted around stem, erect-spreading when moist, obovate, weakly concave, 0.5–2.5 mm;

base not decurrent;

margins recurved to mid leaf, distinctly serrulate distally, limbidium present, of 1–3 rows of cells;

apex acute;

costa long-excurrent, awn sometimes hyaline, slender, irregularly twisted when dry;

proximal laminal cells long-rectangular in rosette leaves;

medial and distal cells short-rhomboidal, 12–25 µm wide, 3–4:1, walls thin, not porose.

of main rosette and innovations similar; slightly twisted to contorted when dry, erect-spreading when moist, broadly ovate to obovate, flat, 1.5–3 mm;

base not decurrent;

margins recurved from mid leaf to near apex, distinctly and strongly serrate distally, limbidium strong, of 2–4 rows of cells;

apex acute;

costa short-excurrent to excurrent, awn stout or slender, irregularly twisted when dry;

proximal laminal cells long-rectangular in rosette leaves;

medial and distal cells long-rhomboidal, 12–20 µm wide, 3–5:1, walls firm, not porose.

Sexual condition

dioicous.

Capsule

inclined, red-brown, cylindric to elongate-pyriform, 3–5 mm.

strongly nutant, red, elongate-pyriform, 3–6 mm.

Specialized

asexual reproduction by rhizoidal tubers, usually on long rhizoids away from stem base, red-brown, same color as rhizoids, 200–300 µm, smooth or almost so.

asexual reproduction by rhizoidal tubers, scarlet, crimson, or red, brighter than rhizoids, (100–)200–300 µm. Sexual condition dioicous or polyoicous.

Rosulabryum capillare

Rosulabryum torquescens

Phenology Capsules mature May–Jul. Capsules mature Apr–Jun.
Habitat Moist shaded soil, soil banks, rotting wood Soil, rock, rotting wood
Elevation low to high elevations (0-2500 m) (low to high elevations (0-8200 ft)) low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft))
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DE; IA; ID; IN; KY; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; Mexico; Central America; South America; Atlantic Islands (including Macaronesia); Pacific Islands (New Zealand); Greenland; Africa; Eurasia; West Indies; Australia
[WildflowerSearch map]
from FNA
AZ; CA; NV; OR; WA; BC; South America; Eurasia; Africa; Atlantic Islands (including Macaronesia); Pacific Islands (New Zealand); Australia
Discussion

Traits diagnostic for the protean Rosulabryum capillare are leaves spirally twisted around the stem, long awn, distinct border, and serrulate margins, small size, rosulate innovations, and rhizoidal tubers the same color as the rhizoids. Rosulabryum gemmascens has irregularly contorted leaves with a much weaker border and nearly entire margins, and elongate somewhat imbricate innovations of small red-brown ovate leaves. Rosulabryum laevifilum has a much more variable costa that sometimes does not reach the apex, green leaves, and filiform gemmae. Rosulabryum torquescens has irregularly contorted leaves with strongly serrate margins and red to crimson tubers, brighter than the rhizoids, and is usually synoicous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rosulabryum torquescens is found on all continents except Antarctica, particularly in seasonal Mediterranean, warm-temperate to subtropical regions, in climates that are warmer and more seasonal than for R. capillare. This is a distinctive species, characterized by irregularly contorted leaves with a short to medium-length awn, strong limbidium, strongly serrate distal margins, mixed sexuality, bright red to crimson tubers, and strongly nutant (at maturity) red capsules. In the flora area, it is most common in Mediterranean climates along the coast, but extends inland in semiarid to arid regions of the North American Southwest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 181. FNA vol. 28, p. 185.
Parent taxa Bryaceae > Rosulabryum Bryaceae > Rosulabryum
Sibling taxa
R. andersonii, R. andicola, R. bornholmense, R. canariense, R. elegans, R. erythroloma, R. flaccidum, R. gemmascens, R. laevifilum, R. pseudocapillare, R. rubens, R. torquescens
R. andersonii, R. andicola, R. bornholmense, R. canariense, R. capillare, R. elegans, R. erythroloma, R. flaccidum, R. gemmascens, R. laevifilum, R. pseudocapillare, R. rubens
Synonyms Bryum capillare Bryum torquescens, B. capillare subsp. torquescens
Name authority (Hedwig) J. R. Spence: Bryologist 99: 223. (1996) (Bruch ex De Notaris) J. R. Spence: Bryologist 99: 223. (1996)
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