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cluster rose, cluster wild rose, peafruit rose, swamp rose

bald-hip rose, dwarf rose, naked-hip rose, wood rose

Habit Shrubs, loosely clustered or in dense thickets. Shrubs or subshrubs, usually loosely clustered.
Stems

ascending to erect, (2–)4–20(–25) dm, openly branched;

bark ± glaucous when young, dark reddish brown or dull red with age outer layer may exfoliate as thin ash gray peel, glabrous;

infrastipular prickles usually paired, erect, rarely curved, usually subulate, 2–10 × 2–4 mm, base glabrous, internodal prickles rare or absent.

erect, sometimes spreading, slender, (1–)3–15(–25) dm, sparsely or densely branched;

bark sometimes glaucous, reddish brown with age, glabrous;

infrastipular prickles 0–2, erect, subulate, 2–8(–10) × 1.5 mm, terete, internodal prickles similar or smaller, sparse to dense, sometimes absent on distal stems, mixed with aciculi, base terete, eglandular.

Leaves

5–10(–13) cm;

stipules 8–22 × 2–5 mm, auricles flared, 2–5 mm, margins usually entire, sometimes erose or lobed, finely ciliolate to ciliate, eglandular, surfaces glabrous or sparsely pubescent, sparsely stipitate-glandular or eglandular;

petiole and rachis sometimes with pricklets, glabrous or hairy hairs to 1 mm, sometimes stipitate-glandular;

leaflets 5–7(–9), terminal: petiolule 8–12 mm, blade elliptic-ovate, (15–)20–45(–60) × 9–16(–20) mm, widest at or below middle, membranous, base cuneate to obtuse, margins 1(–2)-serrate, teeth 12–22 per side, on distal 3/4–4/5 of margin, acute, eglandular, apex acute, sometimes obtuse, abaxial surfaces pale green, usually sparsely pubescent, eglandular, adaxial green, dull, glabrous, rarely puberulent.

(2–)4–10(–17) cm;

stipules 5–15 × 2–5 mm, auricles flared, 2–3 mm, margins entire, stipitate-glandular, surfaces glabrous, eglandular;

petiole and rachis with pricklets, glabrous, rarely finely puberulent, sparsely stipitate-glandular;

leaflets 5–9(–11), terminal: petiolule (2–)5–12(–20) mm, blade elliptic to obovate or ovate to nearly orbiculate, (4–)10–40(–60) × (4–)10–20(–40) mm, membranous to ± leathery, margins 2+-serrate, teeth 7–13 per side, obtuse to acute, gland-tipped, apex obtuse, sometimes nearly acute, rounded, or truncate, abaxial surfaces pale green, glabrous, eglandular, adaxial green, dull, glabrous.

Inflorescences

corymbs, sometimes panicles or solitary flowers, 1–12-flowered.

corymbs, usually 1–3-flowered, rarely in multi-flowered candelabras.

Pedicels

erect, sometimes recurved, slender, 10–22 mm, usually glabrous, sometimes finely puberulent, eglandular, rarely stipulate-glandular;

bracts 2–3, lanceolate, 8–14 × 3–6 mm, margins entire, sometimes serrate, irregularly stipitate-glandular and/or erose, surfaces glabrous or pubescent, eglandular.

erect, ± curved as hips mature, slender, 10–25(–35) mm, glabrous, stipitate-glandular, rarely eglandular (except var. serpentina);

bracts some early caducous, 1 or 2, attached near pedicel bases, ovate or lanceolate, 4–12 × 2–8 mm, margins entire, short stipitate-glandular, apex acute or rounded, surfaces glabrous, eglandular.

Flowers

2.4–3.8 cm diam.;

hypanthium ovoid-urceolate, 3–5 × 2.5–3.5 mm, glabrous, rarely setose, eglandular, neck (0–)0.5–1 × 2 mm;

sepals spreading, ovate-lanceolate, 10–17 × 1.5–3 mm, tip 3–7(–10) × 1.5–2.5 mm, margins entire, abaxial surfaces sometimes puberulent, densely or sparsely stipitate-glandular, rarely eglandular;

petals single, pink to deep pink, 12–18 × 10–18 mm;

stamens 75;

carpels 22–35, styles exsert 1–2 mm beyond stylar orifice (1 mm diam.) of hypanthial disc (3.5 mm diam.).

1.5–3 cm diam.;

hypanthium narrowly ovoid-urceolate, 2–4 × 1.5–2 mm, glabrous, eglandular, neck 0–1 × 1.5 mm;

sepals ascending to reflexed, lanceolate, 5–10 × 2–3 mm, tip 0.1–5 × 1 mm, margins entire, abaxial surfaces glabrous, eglandular, sometimes stipitate-glandular;

petals single, deep pink, 8–15 × 6–13 mm;

stamens 57;

carpels 3–12(–16), styles exsert 1–1.5 mm beyond stylar orifice (1 mm diam.) of hypanthial disc (2–4 mm diam.).

Hips

scarlet, globose, sometimes subglobose or ovoid, 7–15 × 7–13 mm, fleshy, glabrous, eglandular, rarely setose- or stipitate-glandular, neck (0–)1–1.5 × 1.5–3.5 mm;

sepals persistent, erect.

scarlet, irregularly ellipsoid or ellipsoid to nearly globose, 7–15 × 5–13 mm, fleshy, glabrous, eglandular, neck 0–2 × 1.5–2.5 mm;

sepals, styles, and distal receptacle collectively deciduous at fruit maturity along well-defined, circumscissile line, erect to reflexed.

Achenes

basiparietal, 5–35, tan, 3–4 × 1.5–2.5 mm.

basiparietal, (1–)4–10(–12), cream to pale brown, (3–)4.5–7 × 2–4 mm.

2n

= 14, 28.

Rosa pisocarpa

Rosa gymnocarpa

Distribution
from FNA
CA; OR; WA; BC
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[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Rosa gymnocarpa is most easily distinguished in mature fruit, in that the sepals collectively and cleanly separate from the hip together with styles and, sometimes, associated hypanthia. Other diagnostic features include solitary to few, small flowers, stipitate-glandular pedicels, hypanthia extremely small in bud, and glabrous leaflets doubly glandular-toothed. The species is also one of the relatively few roses that flourishes in partial shade. Rosa gymnocarpa occurs in forested areas from British Columbia and Montana to central California, with disjunct populations in southern California; it skirts the Great Basin, with the possible exception of the type of R. leucopsis Greene, a likely synonym, purportedly from central Oregon’s sagebrush steppe, which is otherwise devoid of the species (A. Cronquist and N. H. Holmgren 1997).

The distinctive features of Rosa gymnocarpa have been used as the basis for sect. Gymnocarpae Crépin. Recognition of that section is not supported by the molecular analysis by A. Bruneau et al. (2007), in which R. gymnocarpa occurs in a clade with R. pinetorum and one sample of R. californica, separate from a distinct clade that groups diverse specimens of R. bridgesii, R. spithamea, and some Asian species of sect. Rosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences (1–)3–12-flowered; sepal abaxial surfaces usually stipitate-glandular, tips to 10 mm; infrastipular prickles (1–)2; hips usually globose, rarely subglobose, abruptly narrowed to necks 1.5–3 mm diam.; leaflets most commonly 7, terminal blade 15–35 mm.
subsp. pisocarpa
1. Inflorescences 1–3(–10+)-flowered; sepal abaxial surfaces usually eglandular, rarely stipitate-glandular, tips to 7 mm; infrastipular prickles 0–1(–2); hips subglobose to ovoid, gradually to abruptly narrowed to necks 2.5–3.5 mm diam.; leaflets most commonly 5, terminal blade 20–45(–60) mm.
subsp. ahartii
1. Leaflets (5–)7–9(–11), terminal blades elliptic to narrowly obovate or ovate, (5–)10–30(–60) mm, apices usually obtuse, sometimes nearly acute or rounded; pedicels (10–)15–25(–35) mm; stems 3–15(–25) dm; light shade (rarely full sun) in open woodlands, forest edges, often near streams or rocky sites; s British Columbia to s California, e to Montana.
var. gymnocarpa
1. Leaflets 5(–7), terminal blades broadly elliptic to obovate or ovate to nearly orbiculate, 4–20 mm, apices usually broadly obtuse to rounded, sometimes nearly truncate; pedicels 10–15 mm; stems (1–)3–6(–13) dm; full sun in roadsides, ridges, and other openings in chaparral and stunted forests on ultramafic substrates; Siskiyou Mountains, sw Oregon and nw California.
var. serpentina
Source FNA vol. 9, p. 109. FNA vol. 9, p. 116.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Roseae > Rosa > subg. Rosa > sect. Rosa Rosaceae > subfam. Rosoideae > tribe Roseae > Rosa > subg. Rosa > sect. Rosa
Sibling taxa
R. acicularis, R. arkansana, R. blanda, R. bracteata, R. bridgesii, R. californica, R. canina, R. carolina, R. cinnamomea, R. foliolosa, R. gallica, R. glauca, R. gymnocarpa, R. laevigata, R. lucieae, R. minutifolia, R. mollis, R. multiflora, R. nitida, R. nutkana, R. palustris, R. pinetorum, R. rubiginosa, R. rugosa, R. setigera, R. sherardii, R. spinosissima, R. spithamea, R. stellata, R. tomentosa, R. virginiana, R. woodsii
R. acicularis, R. arkansana, R. blanda, R. bracteata, R. bridgesii, R. californica, R. canina, R. carolina, R. cinnamomea, R. foliolosa, R. gallica, R. glauca, R. laevigata, R. lucieae, R. minutifolia, R. mollis, R. multiflora, R. nitida, R. nutkana, R. palustris, R. pinetorum, R. pisocarpa, R. rubiginosa, R. rugosa, R. setigera, R. sherardii, R. spinosissima, R. spithamea, R. stellata, R. tomentosa, R. virginiana, R. woodsii
Subordinate taxa
R. pisocarpa subsp. ahartii, R. pisocarpa subsp. pisocarpa
R. gymnocarpa var. gymnocarpa, R. gymnocarpa var. serpentina
Name authority A. Gray: Proc. Amer. Acad. Arts 8: 382. (1872) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 461. (1840)
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