Rosa pisocarpa |
Rosa bridgesii |
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cluster rose, cluster wild rose, peafruit rose, swamp rose |
pygmy rose, Sierran dwarf rose, Sierran ground rose, wood rose |
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Habit | Shrubs, loosely clustered or in dense thickets. | Subshrubs, forming open colonies. | ||||
Stems | ascending to erect, (2–)4–20(–25) dm, openly branched; bark ± glaucous when young, dark reddish brown or dull red with age outer layer may exfoliate as thin ash gray peel, glabrous; infrastipular prickles usually paired, erect, rarely curved, usually subulate, 2–10 × 2–4 mm, base glabrous, internodal prickles rare or absent. |
erect, (1–)2–4(–8) dm, openly branched; bark often glaucous when young, gray to dark brown with age, glabrous; infrastipular prickles paired, erect, largest ± flattened, subulate, 3–10 × 2–7 mm, base glabrous, internodal rare or absent, smaller, mixed with aciculi, eglandular. |
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Leaves | 5–10(–13) cm; stipules 8–22 × 2–5 mm, auricles flared, 2–5 mm, margins usually entire, sometimes erose or lobed, finely ciliolate to ciliate, eglandular, surfaces glabrous or sparsely pubescent, sparsely stipitate-glandular or eglandular; petiole and rachis sometimes with pricklets, glabrous or hairy hairs to 1 mm, sometimes stipitate-glandular; leaflets 5–7(–9), terminal: petiolule 8–12 mm, blade elliptic-ovate, (15–)20–45(–60) × 9–16(–20) mm, widest at or below middle, membranous, base cuneate to obtuse, margins 1(–2)-serrate, teeth 12–22 per side, on distal 3/4–4/5 of margin, acute, eglandular, apex acute, sometimes obtuse, abaxial surfaces pale green, usually sparsely pubescent, eglandular, adaxial green, dull, glabrous, rarely puberulent. |
2.5–11 cm; stipules 5–10 × 2–4 mm, auricles flared, 1–3 mm, margins entire, usually sessile-glandular, surfaces glabrous, stipitate-glandular; petiole and rachis with sparse pricklets, glabrous or densely fine hairy hairs to 0.2 mm, densely stipitate-glandular; leaflets (3–)5–7, terminal: petiolule 3–20 mm, blade obovate or widely elliptic-cuneate to ± orbiculate, 10–30(–50) × 8–25(–30) mm, membranous to ± leathery, base rounded-obtuse, margins 2+-crenate, teeth 8–12 per side, obtuse, gland-tipped, apex obtuse to truncate, abaxial surfaces gray-green, finely puberulent, sometimes sparsely stipitate-glandular, adaxial green, dull, finely puberulent to glabrate. |
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Inflorescences | corymbs, sometimes panicles or solitary flowers, 1–12-flowered. |
corymbs, 1 or 2(–7)-flowered. |
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Pedicels | erect, sometimes recurved, slender, 10–22 mm, usually glabrous, sometimes finely puberulent, eglandular, rarely stipulate-glandular; bracts 2–3, lanceolate, 8–14 × 3–6 mm, margins entire, sometimes serrate, irregularly stipitate-glandular and/or erose, surfaces glabrous or pubescent, eglandular. |
erect, relatively slender, 5–17 mm, glabrous, eglandular or stipitate-glandular; bracts 1–3, lanceolate, 3–10 × 1–6 mm, margins entire, stipitate-glandular, surfaces glabrous, eglandular. |
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Flowers | 2.4–3.8 cm diam.; hypanthium ovoid-urceolate, 3–5 × 2.5–3.5 mm, glabrous, rarely setose, eglandular, neck (0–)0.5–1 × 2 mm; sepals spreading, ovate-lanceolate, 10–17 × 1.5–3 mm, tip 3–7(–10) × 1.5–2.5 mm, margins entire, abaxial surfaces sometimes puberulent, densely or sparsely stipitate-glandular, rarely eglandular; petals single, pink to deep pink, 12–18 × 10–18 mm; stamens 75; carpels 22–35, styles exsert 1–2 mm beyond stylar orifice (1 mm diam.) of hypanthial disc (3.5 mm diam.). |
2.5–3.5 cm diam.; hypanthium ovoid-globose to widely urceolate, 4–5 × 3–4 mm, glabrous, rarely sparsely setose, usually eglandular, neck 0.5–1(–2) × 1.5–3 mm; sepals spreading, lanceolate, 6–14 × 2–3 mm, tip 0–4 × 0.5 mm, margins entire, abaxial surfaces puberulent, usually stipitate-glandular; petals single, deep pink, 10–20 × 10–20 mm; carpels 10–30, styles exsert 0.5–1 mm beyond stylar orifice (1–1.5 mm diam.) of hypanthial disc (2–4 mm diam.). |
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Hips | scarlet, globose, sometimes subglobose or ovoid, 7–15 × 7–13 mm, fleshy, glabrous, eglandular, rarely setose- or stipitate-glandular, neck (0–)1–1.5 × 1.5–3.5 mm; sepals persistent, erect. |
scarlet, ovoid to depressed-globose to pyriform, (8–)10–18 × 7–14 mm, fleshy, glabrous, rarely sparsely setose distally, eglandular, neck 1 × 2.5–4.5 mm; sepals persistent, erect. |
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Achenes | basiparietal, 5–35, tan, 3–4 × 1.5–2.5 mm. |
basiparietal, 1–11, cream to pale brown, 3.5–5(–6.5) × 2.5–4 mm. |
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2n | = 14, 28. |
= 14, 28. |
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Rosa pisocarpa |
Rosa bridgesii |
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Phenology | Flowering May–Aug. | |||||
Habitat | Open forest floors, meadow edges, rocky outcrops, midmontane forests, shade to sun | |||||
Elevation | 700–2500 m (2300–8200 ft) | |||||
Distribution |
CA; OR; WA; BC
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CA; OR
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As here circumscribed, Rosa bridgesii is the Sierran-Cascade counterpart of R. spithamea, encompassing previous references to R. pinetorum and R. spithamea from the Sierra Nevada and most applications of R. yainacensis. The species is the low-growing, openly rhizomatous rose of midmontane forests in the Sierra Nevada and southern Cascade Range of California and Oregon. Finely puberulent leaflets help distinguish non-fruiting R. bridgesii from sympatric R. gymnocarpa, which has glabrous leaflets. Plants from Oregon differ from Sierran plants in that prickles are less likely to be infrastipular only. In the phylogenetic analysis by A. Bruneau et al. (2007), Rosa bridgesii and R. spithamea comprise a distinct clade more closely related to Asian species than to others from western North America. Rosa bridgesii has been conserved against R. calvaria Greene, R. covillei Greene, R. crenulata Greene, R. myriadenia Greene, and R. yainacensis Greene (B. Ertter 2007b). Only R. calvaria and R. crenulata are unambiguous synonyms of R. bridgesii; the taxonomic identity of the other names remains unresolved and may involve hybridization with other species. On the basis of pollen size, E. W. Erlanson (1931) believed that material falling within the current circumscription of Rosa bridgesii was tetraploid (2n = 28). In a subsequent paper (1934), she reported a diploid somatic count (2n = 14) for R. calvaria, a synonym of R. bridgesii. Recent flow cytometry indicates a tetraploid condition for the sampled specimen of R. bridgesii (A. Bruneau et al., unpubl. data). It is possible that both ploidy levels are present within the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 109. | FNA vol. 9, p. 118. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Roseae > Rosa > subg. Rosa > sect. Rosa | Rosaceae > subfam. Rosoideae > tribe Roseae > Rosa > subg. Rosa > sect. Rosa | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | R. gymnocarpa var. pubescens, R. spithamea var. solitaria, R. spithamea var. subinermis | |||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 8: 382. (1872) | Crépin ex Rydberg: Bull. Torrey Bot. Club 44: 83. (1917) | ||||
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