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knieskern's beaksedge

Harvey's beaksedge

Habit Plants perennial, cespitose, to 50 cm; rhizomes absent. Plants perennial, cespitose, 70–110 cm; rhizomes absent.
Culms

erect to arching, leafy, linear to filiform, nearly triangular.

erect to excurved, leafy, obscurely trigonous, slender.

Leaves

ascending, overtopped by culm;

blades flat, linear to filiform, to 1.8 mm wide, apex distally involute, trigonous, setaceous.

spreading to ascending, shorter than culm, crowded toward culm base;

blades linear, proximally flat, 1–3 mm wide, gradually involute, apically trigonous, subulate.

Inflorescences

terminal and axillary, spikelet clusters 2–4, widely spaced, the lowest near plant base;

clusters compact, broadly turbinate to hemispheric, to 1.5 cm wide; leafy bracts curved, setaceous, slightly to greatly overtopping subtended compounds.

spikelet clusters 1–4, dense to open, mostly irregularly turbinate;

peduncles ascending, branches spreading to erect, ultimate branches with many spikelets; leafy bracts setaceoustipped, usually exceeding all clusters, or at least all but the distal.

Spikelets

dark brown, lance-ellipsoid, 2–3 mm;

fertile scales 2 mm, apex acute, midrib short-excurrent or not.

light redbrown or brown, broadly ellipsoid to lanceoloid, 3–4 mm, apex acute to acuminate;

fertile scales ovate to obovate or suborbiculate, 2–3.5 mm apex acute to rounded or emarginate, midrib included or exserted as mucro.

Flowers

perianth bristles 6, ± as long as fruit body, retrorsely barbellate.

perianth bristles mostly 6, rarely reaching fruit midbody, antrorsely barbellate.

Fruits

mostly 2 per spikelet, 1.5–1.9 mm;

body brown with yellowish center, ellipsoid, lenticular distal to short stipe, 1–1.3 × 0.6–0.8 mm;

tubercle triangular, 0.3–0.6 mm, distinctly shorter than fruit body.

mostly 1 per spikelet, 2–2.5 mm, body dark brown, obovoid to subglobose, tumid or lenticular, 1.5–1.7 mm, transversely finely rugose to nearly level, intervals with very small, pitlike alveoli.

Rhynchospora knieskernii

Rhynchospora harveyi

Phenology Fruiting summer–fall.
Habitat Moist to wet pine barrens, sand pits, borrow pits
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
DE; NJ
[BONAP county map]
from FNA
AL; AR; FL; GA; KS; LA; MO; MS; NC; OK; SC; TN; TX; VA; se United States; Midwestern
[BONAP county map]
Discussion

Of conservation concern.

Rhynchospora knieskernii is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit body broadly obovoid to suborbicular, medially with mostly isodiametric tiny alveoli or pits, or minutely raised reticulate in an almost honeycomb pattern of alveolae, or simply evenly finely cancellate; ultimate spikelet complexes with clusters on stiffish branchlets, usually dense and exceeded at least by subulate tips of subtending leafy bract and bractlets.
var. harveyi
1. Fruit body obovoid, lenticular, medially with oblong or roundish pitlike alveoli, intervals between contiguous transverse rows forming shallow, broad, pale, smooth ridges; ultimate spikelet clusters more sparse, on more slender, lax, erect to excurved branches and exceeding subtending bracts and bractlets.
var. culixa
Source FNA vol. 23, p. 211. FNA vol. 23, p. 231.
Parent taxa Cyperaceae > Rhynchospora Cyperaceae > Rhynchospora
Sibling taxa
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. harveyi, R. indianolensis, R. inexpansa, R. inundata, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
Subordinate taxa
R. harveyi var. culixa, R. harveyi var. harveyi
Name authority J. Carey: Amer. J. Sci. Arts, ser. 2, 4: 25. (1847) W. Boott: Bot. Gaz. 9: 85. (1884)
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