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Harvey's beaksedge

pineland beaksedge

Habit Plants perennial, cespitose, 70–110 cm; rhizomes absent. Plants perennial, cespitose, 50–110 cm; rhizomes absent.
Culms

erect to excurved, leafy, obscurely trigonous, slender.

lax, often excurved, slender, ± terete or trigonous.

Leaves

spreading to ascending, shorter than culm, crowded toward culm base;

blades linear, proximally flat, 1–3 mm wide, gradually involute, apically trigonous, subulate.

ascending, exceeded by culm;

blades linear, proximally flat, 1.5–2.5 mm wide, apex trigonous, subulate, tapering.

Inflorescences

spikelet clusters 1–4, dense to open, mostly irregularly turbinate;

peduncles ascending, branches spreading to erect, ultimate branches with many spikelets; leafy bracts setaceoustipped, usually exceeding all clusters, or at least all but the distal.

clusters 3–4, widely spaced, narrowly, compactly, or diffusely turbinate; leafy bracts exceeding proximal clusters.

Spikelets

light redbrown or brown, broadly ellipsoid to lanceoloid, 3–4 mm, apex acute to acuminate;

fertile scales ovate to obovate or suborbiculate, 2–3.5 mm apex acute to rounded or emarginate, midrib included or exserted as mucro.

deep redbrown, ovoid to broadly ellipsoid, 2–3 mm, apex acute to acuminate;

fertile scales broadly elliptic to obovate or orbiculate, 1.4–2(–2.5) mm, apex rounded to notched, midrib shortexcurrent.

Flowers

perianth bristles mostly 6, rarely reaching fruit midbody, antrorsely barbellate.

perianth bristles 0–3, vestigial when present.

Fruits

mostly 1 per spikelet, 2–2.5 mm, body dark brown, obovoid to subglobose, tumid or lenticular, 1.5–1.7 mm, transversely finely rugose to nearly level, intervals with very small, pitlike alveoli.

2–4 per spikelet, 1.5 mm;

body pale brown to brown, strongly flattened, orbicular to broadly obovoid, 1–1.3 × 0.9–1.2 mm, surfaces sharply transversely wavyrugose, intervals finely vertically striate with rows of linearrectangular alveolae;

tubercle depressed, triangular, flattened, 0.2–0.3 mm, base lunate.

Rhynchospora harveyi

Rhynchospora perplexa

Phenology Fruiting late spring–fall or all year (south).
Habitat Sands and peats of pond and lakeshores, depressions in savannas and flatwoods, or seeps
Elevation 0–400 m (0–1300 ft)
Distribution
from FNA
AL; AR; FL; GA; KS; LA; MO; MS; NC; OK; SC; TN; TX; VA; se United States; Midwestern
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; NC; SC; TN; TX; VA; Central America; West Indies (Cuba, Dominican Republic)
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In habit and in shape, size, and color of spikelet, Rhynchospora perplexa strongly resembles R. microcarpa, a species with which it is commonly associated in the Coastal Plain. An examination of the fruit shows those of R. perplexa to be flattened, with fewer and much coarser transverse ridges, the intervals with very narrow vertical alveolae. The perianth in most instances is absent or rudimentary. Fruit of R. microcarpa is biconvex with more transverse ridges (eight or more), the intervals more coarsely alveolate; its perianth bristles are six, evident, extending at least halfway up the fruit body.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit body broadly obovoid to suborbicular, medially with mostly isodiametric tiny alveoli or pits, or minutely raised reticulate in an almost honeycomb pattern of alveolae, or simply evenly finely cancellate; ultimate spikelet complexes with clusters on stiffish branchlets, usually dense and exceeded at least by subulate tips of subtending leafy bract and bractlets.
var. harveyi
1. Fruit body obovoid, lenticular, medially with oblong or roundish pitlike alveoli, intervals between contiguous transverse rows forming shallow, broad, pale, smooth ridges; ultimate spikelet clusters more sparse, on more slender, lax, erect to excurved branches and exceeding subtending bracts and bractlets.
var. culixa
Source FNA vol. 23, p. 231. FNA vol. 23, p. 229.
Parent taxa Cyperaceae > Rhynchospora Cyperaceae > Rhynchospora
Sibling taxa
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. harveyi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
Subordinate taxa
R. harveyi var. culixa, R. harveyi var. harveyi
Synonyms Phaeocephalum perplexum
Name authority W. Boott: Bot. Gaz. 9: 85. (1884) Britton: in J. K. Small, Fl. S.E. U.S., 197, 1328. (1903)
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