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Harvey's beaksedge

brownish beak-rush, brownish beaksedge, capitate beak rush, rhynchospore à petites têtes

Habit Plants perennial, cespitose, 70–110 cm; rhizomes absent. Plants perennial, cespitose, 20–100 cm; rhizomes absent.
Culms

erect to excurved, leafy, obscurely trigonous, slender.

arching-ascending, leafy, obtusely trigonous, slender;

principal leaves overtopped by inflorescence;

blades flat, to 3 mm, apex tapering, trigonous.

Leaves

spreading to ascending, shorter than culm, crowded toward culm base;

blades linear, proximally flat, 1–3 mm wide, gradually involute, apically trigonous, subulate.

Inflorescences

spikelet clusters 1–4, dense to open, mostly irregularly turbinate;

peduncles ascending, branches spreading to erect, ultimate branches with many spikelets; leafy bracts setaceoustipped, usually exceeding all clusters, or at least all but the distal.

terminal and axillary, clusters 1–5 or more, compact, turbinate or hemispheric, 1–1.5 cm wide;

peduncles progressively shorter distally on culm;

bracteal leaves mostly exceeding subtended compounds.

Spikelets

light redbrown or brown, broadly ellipsoid to lanceoloid, 3–4 mm, apex acute to acuminate;

fertile scales ovate to obovate or suborbiculate, 2–3.5 mm apex acute to rounded or emarginate, midrib included or exserted as mucro.

rich deep brown, rarely pale brown, lanceellipsoid, 3.5–4(–5) mm;

fertile scale elliptic, 2.7–3 mm, apex acute or rounded, midrib shortexcurrent or not.

Flowers

perianth bristles mostly 6, rarely reaching fruit midbody, antrorsely barbellate.

perianth absent.

Fruits

mostly 1 per spikelet, 2–2.5 mm, body dark brown, obovoid to subglobose, tumid or lenticular, 1.5–1.7 mm, transversely finely rugose to nearly level, intervals with very small, pitlike alveoli.

(1–)2–3(–5) per spikelet, (2–)2.5–3 mm;

body redbrown with pale central disc, stipitate, lenticular, obovoid, 1.2–1.5 × 0.7–1(–1.2) mm, margins pale, wirelike, surfaces slick;

tubercle triangular-subulate, (0.8–)1–1.2(–1.6) mm.

Rhynchospora harveyi

Rhynchospora capitellata

Phenology Fruiting summer–fall.
Habitat Moist to wet meadows, swales, seeps, stream banks, flatwoods, fens, and bogs
Elevation 0–1600 m (0–5200 ft)
Distribution
from FNA
AL; AR; FL; GA; KS; LA; MO; MS; NC; OK; SC; TN; TX; VA; se United States; Midwestern
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; OR; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Rhynchospora capitellata occurs infrequently in the lower Gulf coastal plain (in Florida, only north) and intergrades with R. glomerata. Forms with antrorsely barbellate bristles are referred to forma controversa (S. F. Blake) Gale; those with smooth bristles are named forma discutiens (C. B. Clarke) Gale (S. Gale 1944). Occasional forms with trigonous fruits occur.

Bruce Sorrie (pers. comm.) believes that many of the southern coastal plain records for Rhynchospora capitellata are based on examples of a neglected taxon, R. leptocarpa (Chapman) Small (J. K. Small 1933), and he wishes to reinstate it. Occurring in semi-shady moist sites in steepheads, or from seeps and shallows along blackwater streams, the plants are mostly distinguished by the tall, lax habit; the many distant, small clusters of light brown or tan spikelets; and the softer, more lax foliage. Measures of spikelets, fertile scales, perianth, achenes, and fruit tubercles show a strong overlap with those of southern examples of R. capitellata. The long, lax culm habit, the softer and paler foliage, and the paler spikelets could well be ecologic responses, or the plants could indeed represent a geographic variant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit body broadly obovoid to suborbicular, medially with mostly isodiametric tiny alveoli or pits, or minutely raised reticulate in an almost honeycomb pattern of alveolae, or simply evenly finely cancellate; ultimate spikelet complexes with clusters on stiffish branchlets, usually dense and exceeded at least by subulate tips of subtending leafy bract and bractlets.
var. harveyi
1. Fruit body obovoid, lenticular, medially with oblong or roundish pitlike alveoli, intervals between contiguous transverse rows forming shallow, broad, pale, smooth ridges; ultimate spikelet clusters more sparse, on more slender, lax, erect to excurved branches and exceeding subtending bracts and bractlets.
var. culixa
Source FNA vol. 23, p. 231. FNA vol. 23, p. 210.
Parent taxa Cyperaceae > Rhynchospora Cyperaceae > Rhynchospora
Sibling taxa
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. harveyi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
Subordinate taxa
R. harveyi var. culixa, R. harveyi var. harveyi
Synonyms Schoenus capitellatus, Phaeocephalum glomeratum var. minus, R. capitellata var. leptocarpa, R. capitellata var. minor, R. glomerata var. minor, R. leptocarpa
Name authority W. Boott: Bot. Gaz. 9: 85. (1884) (Michaux) Vahl: Enum. Pl. 2: 235. (1805)
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