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anglestem beaksedge

Harvey's beaksedge

Habit Plants perennial, cespitose, 70–150 cm; rhizomes often present, short, scaly. Plants perennial, cespitose, 70–110 cm; rhizomes absent.
Culms

erect or ascending, leafy, trigonous.

erect to excurved, leafy, obscurely trigonous, slender.

Leaves

exceeded by culm;

blades linear, proximally 4–7 mm wide, apex trigonous, tapering.

spreading to ascending, shorter than culm, crowded toward culm base;

blades linear, proximally flat, 1–3 mm wide, gradually involute, apically trigonous, subulate.

Inflorescences

terminal and axillary;

clusters 3–6, mostly dense, narrowly to broadly turbinate, branches ascending; leafy bracts exceeeding proximalmost inflorescences.

spikelet clusters 1–4, dense to open, mostly irregularly turbinate;

peduncles ascending, branches spreading to erect, ultimate branches with many spikelets; leafy bracts setaceoustipped, usually exceeding all clusters, or at least all but the distal.

Spikelets

rich brown, ovoid, (3–)4–5 mm, apex acuminate;

fertile scales ovate, 2.5–3.5 mm, apex acuminate, midrib included or shortexcurrent.

light redbrown or brown, broadly ellipsoid to lanceoloid, 3–4 mm, apex acute to acuminate;

fertile scales ovate to obovate or suborbiculate, 2–3.5 mm apex acute to rounded or emarginate, midrib included or exserted as mucro.

Flowers

perianth bristles mostly 6, exceeding tubercle tip.

perianth bristles mostly 6, rarely reaching fruit midbody, antrorsely barbellate.

Fruits

mostly 3–4 per spikelet, 2–2.2 mm;

body brown on short pedicellar (to 0.3 mm) stalk, broadly obovoid, lenticular, 1.3–1.5 × 1–1.5 mm, surfaces transversely rugulose, vertically finely striate and rectangularalveolate;

tubercle compressed, triangular acuminate, 0.5–0.8 mm, edges setulose.

mostly 1 per spikelet, 2–2.5 mm, body dark brown, obovoid to subglobose, tumid or lenticular, 1.5–1.7 mm, transversely finely rugose to nearly level, intervals with very small, pitlike alveoli.

Rhynchospora caduca

Rhynchospora harveyi

Phenology Fruiting summer–fall.
Habitat Low meadows, clearings, marshes, marsh borders, seeps, bog moats, savannas, ditches, pine flatwoods, swamps
Elevation 0–400 m (0–1300 ft)
Distribution
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; KS; LA; MO; MS; NC; OK; SC; TN; TX; VA; se United States; Midwestern
[BONAP county map]
Discussion

Rhynchospora caduca has its closest relationships with the even more robust R. odorata Grisebach, on the one hand, and the swampinhabiting, more slender, and rhizomatous R. mixta Britton ex Small, on the other. Intergrades with R. odorata appear in Alabama and northwest Florida; intergrades with R. mixta appear where ranges overlap in both the Atlantic and Gulf coastal plains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit body broadly obovoid to suborbicular, medially with mostly isodiametric tiny alveoli or pits, or minutely raised reticulate in an almost honeycomb pattern of alveolae, or simply evenly finely cancellate; ultimate spikelet complexes with clusters on stiffish branchlets, usually dense and exceeded at least by subulate tips of subtending leafy bract and bractlets.
var. harveyi
1. Fruit body obovoid, lenticular, medially with oblong or roundish pitlike alveoli, intervals between contiguous transverse rows forming shallow, broad, pale, smooth ridges; ultimate spikelet clusters more sparse, on more slender, lax, erect to excurved branches and exceeding subtending bracts and bractlets.
var. culixa
Source FNA vol. 23, p. 223. FNA vol. 23, p. 231.
Parent taxa Cyperaceae > Rhynchospora Cyperaceae > Rhynchospora
Sibling taxa
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. harveyi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
R. alba, R. baldwinii, R. brachychaeta, R. breviseta, R. caduca, R. californica, R. capillacea, R. capitellata, R. careyana, R. cephalantha, R. chalarocephala, R. chapmanii, R. ciliaris, R. colorata, R. compressa, R. corniculata, R. crinipes, R. curtissii, R. debilis, R. decurrens, R. divergens, R. elliottii, R. eximia, R. fascicularis, R. fernaldii, R. filifolia, R. floridensis, R. fusca, R. globularis, R. glomerata, R. gracilenta, R. grayi, R. harperi, R. indianolensis, R. inexpansa, R. inundata, R. knieskernii, R. kunthii, R. latifolia, R. macra, R. macrostachya, R. megalocarpa, R. megaplumosa, R. microcarpa, R. microcephala, R. miliacea, R. mixta, R. nitens, R. nivea, R. odorata, R. oligantha, R. pallida, R. perplexa, R. pineticola, R. pleiantha, R. plumosa, R. punctata, R. pusilla, R. rariflora, R. recognita, R. scirpoides, R. solitaria, R. stenophylla, R. thornei, R. torreyana, R. tracyi, R. wrightiana
Subordinate taxa
R. harveyi var. culixa, R. harveyi var. harveyi
Synonyms Phaeocephalum caducum, R. patula
Name authority Elliott: Sketch Bot. S. Carolina 1: 62. (1816) W. Boott: Bot. Gaz. 9: 85. (1884)
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